Diplocladus louvelii ( Spinola, 1844 )

Gerstmeier, Roland & Weiss, Ingmar, 2009, Revision of the Genera Diplocladus Fairmaire and Strotocera Schenkling (Coleoptera: Cleridae, Tillinae), Zootaxa 2242, pp. 1-54 : 14-16

Diplocladus louvelii ( Spinola, 1844)

(Figs. 8, 45–46)

Callitheres louvelii Spinola, 1844: 111 ; Desmarest 1860: 236, fig. 163; Schenkling 1902: 123; Schenkling 1906: 245 ( Strotocera ); Heyne & Taschenberg 1908: plate 26, fig. 47 ( Pallenis ); Lesne 1922: 653 ( Diplocladus ).

Cymatodera aschantina Kuwert, 1893: 468 ; Kraatz 1899: 86 ( Pallenis ); Schenkling 1899: 334 ( Eucymatodera ); Schenkling 1902: 123 ( Strotocera ).

Strotocera subnigra Pic, 1949 ; Ekis 1975: 11.

Diplocladus tilloides Fairmaire, 1887: 161 ; Lesne 1922: 653, plate I c-10, fig. 6, ( Diplophorus ) n. syn. Diplophorus ? pectorali s Hintz, 1902, syn.

Diplocladus rhodesianus Péringuey, 1889: 242 (Philocalus) ; Schenkling 1906: 244; Lesne 1922: 653 ( Diplophorus ) n. syn.

Specimens examined:

Lectotype: Diplocladus louvelii (Spinola) (MRSN): Ƥ. Ekis 1974.

Paralectotype: Diplocladus louvelii (Spinola) (MRSN): Ƥ. Ekis 1974.

Holotype: D. O. Afrika; E.Fischer S, 3, 76571, Type, Pallenis pectoralis, HOLOTYPUS, Pallenis pectoralis, Hintz 1902, labelled by MINHUB 2008 (Ƥ, ZMB); Diplocladus tilloides n. syn.

Holotype: Rhodesia, Buluwayo, Selous, Philocalus rhodesianus , typ, PJ, Type, SAM Type, Acc. no. 1270 (Ƥ, SAMC); Diplocladus rhodesianus n. syn.

Holotype: Goldküste, Ex. Musaeo, A. Kuwert, 1894, aschantina , kuw. typ!, TYPE (2 ƤƤ, MNHN); = Cymatodera aschantina Kuwert, 1893 , syn.

Holotype: Togo, Bismarckburg, 25.IX.–12.XI.92, L. Conradt S., pectoralis, Hintz, Determ. E. Hintz., Ex. Musoeo, E. Hintz, Museum Paris,1952, Coll R Oberthur, TYPE (Ƥ, MNHN); = Diplophorus ? pectoralis Hintz, 1902, syn.

Others: 27.v.1944, E. Africa, Tang T’y., Lohnmbo, Tinde d.d., E. Burtt., 33.1948, 27/5/44, one seen, Sterotocera louveli Spinola, E.A.J. Duffy det. 1948 (1 ex., HEC); Kenya: central south, Kibwezi Forest, Spring Camp 28.–29.11., 2004 leg. Puchner (2 ex., CRS); Zoutpan, Pta, 4–10/II 1929, G.v. Son (1 ex., TMSA); UNREADABLE, 5–28 (1 ex., TMSA); Abachaus SWA, II 1944, G. Hobohm (1 ex., TMSA); Abachaus SWA, III 1944, G. Hobohm (1 ex., TMSA); Msali Bridge, 17–19.IV.1954, A.J.T. Janse, Diplocladus louvelii, (Spinola) , det. G. Ekis 1985 (1 ex., TMSA); UNREADABLE, 4-II-28, UNREADABLE ( TMSA); Abachaus, S.W.A, May 44, G. Hobohm (1 ex., TMSA); Mozambique, SofalaP, 30km S Caia, 76m, 18.02S - 35.12E, 21.12.2005; E-Y 3709, light trap, leg. Gussmann, Müller (1 ex., TMSA); Zoutpan, Pta, 4–10/II 1929, G. v. Son, rhodesiana, Rh. Peringuey (1 ex., TMSA); Central Malawi, 5 km W, Golomoti S 14.25 E 43.33, 22– 23.I.1985 650m, CL Bellamy, S Chown, AV Evans, AS Schoeman, CH Scholtz (1 ex., SANC); Central Malawi, North Senga, Forest Res S 13.43 E 34.37, 23–26.I.1985 480m, CL Bellamy, S Chown, AV Evans, AS Schoeman, CH Scholtz, National Coll. of Insects, Pretoria, South Africa (1 ex., SANC); South Africa, Transvaal, Kruger Nat. Park, Punda Maria, 22.41S 31.02E, 07.ii.1988, G.D. Butler (1 ex., SANC); South Africa, Transvaal, Weltevreden Farm, nr Nelspruit, 25.34S 31.10E, 02.ii.1989, R. Oberprieler (1 ex., SANC); South Africa, Transvaal, Kralingen, 35 km, NW Warmbad. 24.37S, 28.06E, 17.i.1983, M.W. Mansell (1 ex., SANC); South Africa: KwaZulu-Natal, Iphiva Camp., St Lucia Game Reserve, ca. 28.05S 32.20E, 11.ii.1997 R. Oberprieler (1 ex., SANC); South Africa, Natal, Lake Cubhu, 28.52S, 31.58E iii.1982, Light trap, R. Oberprieler (1 ex., SANC); South Africa, Natal, Empangeni, 28.44S, 31.54E 26.iii., 1990 P.E. Reavell (1 ex., SANC); South Africa, Natal, Empangeni, 28.44S, 31.54E ii. 1985 P. Reavell (1 ex., SANC); South Africa, Transvaal, Vienna Game Farm, near Hoedspruit, 24.17S 30.58E, 17–18.i.1991, E. Grobbelaar, Collected at light (2 ex., SANC); South Africa, Transvaal, Kruger Nat. Park, Renosterkoppies, 25.07S 31.36E, 450m. 17.i.1985, N.C. Piennaar, Collected at light (2 ex., SANC); South Africa: MPU, Mahushe Shongwe N.R., near Mzinti 25.45S, 31.44E, 04–05.ii.1995, Chown, Stals, Browne (1 ex., SANC); South Africa, Kwazulu, Mseleni, Miss. 27.23S 32.32E, iii.1985 P. Reavell, light trap (1 ex., SANC); South Africa, Natal, Mtunzini, 28.58S, 31.46E iii.1989, P. Reavell, Light trap (1 ex., SANC); Lamto, Toumodi, Côte d’Ivoire, 14. VIII. 1971, Col. Chemin ( MNHN); Guinea (2 ex., NHMB). Republik Guinea, Sérédou, 9. Nov. 1974, leg. Dr. Zott (1 ex., ZMB); Hist.-Coll. ( Coleoptera ), Nr. 17799, Phloiocopus louvelii Spin., Guineam Deyrolle, Zool. Mus. Berlin (1 ex., ZMB); 17799, Hist.-Coll. ( Coleoptera ), Nr. 17799, Phloiocopus louvelii Spin., Guineam Deyrolle, Zool. Mus. Berlin, Louvelii, Spin. , Guinea Deyr. (1 ex., ZMB); 367, Togo, Bismarckburg, L. Conradt S., Strotocera pectoralis, Hintz, 19, Schenkling det. (1 ex., ZMB); Afr.or., Luitpoldkette, Strotocera pectoralis, Hintz, Mit Type verglichen (1 ex., ZMB); Afr.or. Kitwi (1 ex., ZMB); Togo, Bismarckburg, 3.–10.XII.92, L. Conradt S., 66 (1 ex., ZMB); Südafrika: Transvaal, Blyderivierspoort Nat Res, 24°39’S / 30°50’E, 13.xii.1995, leg. F Koch (1 ex., ZMB); Togo, Misahöhe, 12.X.93, E. Baumann S., C272, Misahöhe 12.X.93, leg. Baumann, fliegt des Abends (1 ex., ZMB); Togo, Conradt, Coll. Kraatz, Kraatz det. (1 ex. DEI); Togo, Cymatodera Aschantia, Belg. 93468, Kuwert, Pallenis Louvelii, Pic , coll. Kraatz, Kraatz det., coll. DEI Müncheberg, Strotocera louvelii, Spin. (1 ex., DEI); UNREADABLE im Togogebiet, Staudinger & Bang- Haas dedic., Schenkling det. (1 ex., DEI); Nigeria, Takura, 13p State 19 Aug 1973, Col. J.T. Medler, CX-46 (1 ex., MRAC); Musée du Congo, Dr Sheffield Neave, Kayambo-Dikulwe VI.07, Strotocera louvelii Spin., R. DÉT., B, 377 (1 ex., MRAC); Musée du Congo, Mahagi-Niarembe, - V-1935, Ch. Scops, R. DET., C, 5490, Strotocera pectoralis, Hintz (1 ex., MRAC); Port.E.Africa., Valley of Kola R, nr.E.Mt.Chiperone, 1, 500- 2,000ft.,3 Apl.1913, S.A. Neave, 1913–140, Diplocl. rhodesianus (1 ex., BMNH); 6696, Salisburry, Mashonaland, Jan. 1895, G.A.K. Marshall, Marshall Coll., 1914–554, Strotocera rhodesiana (Per) (1 ex., BMNH); Kano Dist, N. Nigeria, aug 1954, W.E.S. Merrett, B.M.1962-270, Diplocladus, Ginter Ekis det. 19 (1 ex., BMNH); MAS Arthur, Sambu Masai 1 35, Imp.Inst, Entom., Pres by Com Inst Ent, B.M. 1965-2 (1 ex., BMNH); Tanganyika, Old Shinyanga., 14.II.54, E. Burtt., UNREADABLE, Pres by Com Inst Ent, B.M. 1955-292, larves de cette UNREADABLE colbee dans combratum abocatum, ds galenis de Salagena sp., ref.82.159 (1 ex., BMNH); Obervolta, Folonzo, XI.89, H. Politzar (CRG); Obervolta, Bobo Diulasso, XI.81, H. Politzar (2 ex., CRG); Obervolta, Bobo Diulasso, X.82, H. Politzar (2 ex., CRG); Ober-Volta, Folonzo, III.85, H. Politzar, Tillus spec. (1 ex., CRG); Ober-Volta, Folonzo, III.85, H. Politzar (1 ex., CRG); Obervolta, Bobo Diulasso, 9.1985, leg. H. Politzar (1 ex., CRG); Nigeria /Afr., Molkwa, 26X.1970, Politzar leg. (1 ex., CRG); Kamerun, 6.1991, Nachtigal (7 ex., CRG); Kenya mer., 50km N of Namanga, Ilbisil env., 18.xi.1997, lgt. M. Snizek (1 ex., CRG); Tanzania, I 1990, Kilimanjaro Prov., Same Pare Mts, 1000m leg. Werner (1 ex., CRG), Kankiye, 12.6.1965 (1 ex., CRG); Tanzania, Iringa Dist., Pavvanga near Itunundu/Kisanga/Ruaha NP 800m, 6.1.1993 leg. Mourglia (1 ex., CRG); Kenya - Meru Distr., Materi (Mitunguu), mt. 800-, R. Mourglia legit, 5/ 13.11.1988 (1 ex., CRG); Kenya-S, Voi, 23.xi.1997, M. Snizek leg. (1 ex., CRG); Kenya,Voi, Sagala Reg. 12.1991, leg. K.Werner (1 ex., CRG); Kenya,Voi, Sagala Region, 11.1992 leg. Werner (1 ex., CRG); Guinea, D. UNREADABLE (1 ex., MRSN).

Length: 7.1mm – 19.4mm (80 specimens).

Head: Black, densely and finely punctate.

Antennae: Black; almost reaching base of pronotum when laid alongside.

Pronotum: Dark brown to black, lateral margins and base reddish; anterior half parallel, only slightly broader in middle, constricted posteriorly; surface anterior to transverse depression finely and diffusely punctate, coarsely and irregularly wrinkled behind transverse depression; disc with two patches of white to yellowish hairs; length:width ratio 1.49:1.

Scutellum: Dark red-brown to black.

Elytra: Basal third to half reddish brown, remainder black with white transverse fasciae beyond middle spanning from lateral to sutural margins (not always reaching the suture); disc regularly punctate (punctures often elongate), at shoulders diameter of punctures smaller than interstices, behind shoulders more or less equal, sometimes slightly larger, punctures within fasciae small and sparsely distributed, diameter of punctures strongly variable behind fasciae (punctures mostly elongate); length:width ratio 2.51:1. Legs: Dark brown to black.

Lower surface: Head and thorax red-brown, abdomen black.

Vestiture: With whitish to yellowish hairs, pronotal margins vested with white, disc with reddish hairs, elytra with white hairs, with isolated shorter black hairs.

Distribution: Burkina Faso, Cameroon, Congo, Ghana, Guinea, Ivory Coast, Kenya, Malawi, Mozambique, Namibia, Nigeria, Senegal, Somalia, South Africa, Sudan, Tanzania, Togo, Zimbabwe.