Nebria (Epinebriola) triseriata, C.Huber & J.Schmidt, 2017

Huber, Charles & Schmidt, Joachim, 2017, Notes on the Nebria subgenus Epinebriola K. DANIEL, 1904 with the description of Barbonebriola subgen. nov. and 13 new species from the Himalaya-Tibet orogen (Coleoptera, Carabidae, Nebriini), Contributions to Natural History 36, pp. 1-85 : 35-37

publication ID	https://doi.org/ 10.5169/seals-787049
DOI	https://doi.org/10.5281/zenodo.6362817
persistent identifier	https://treatment.plazi.org/id/03948799-FFF9-FFE4-FF41-FEE2FE2FF9C3
treatment provided by	Carolina
scientific name	Nebria (Epinebriola) triseriata
status	sp. nov.

Treatment

Nebria (Epinebriola) triseriata View in CoL sp. nov. ( Fig. 19 View Fig )

Holotype ♂: Nepal, Dhaulagiri Zone, Myagdi Distr., Annapurna South Himal , W-Slope , 28° 28,30' N, 83° 44,55' E, 4500 m, 22. 5. 2001, leg. G. Hirthe & J. Schmidt ( cSCHM). GoogleMaps

Paratypes: 17 ♂ 13 ♀, same data as holotype ( NMBE, cSCHM) GoogleMaps .

Body length 11–12 mm. Colour black. Pronotum dark brown, somewhat lighter than the head. Mandibulae, appendages of head and antennae brown. Legs black, tarsi brown.

Head with a shallow transverse impression behind the prominent eyes. Mandibulae long. Anterior margin of the labrum distinctly convex and bulging, bearing six setae. Apical margin of clypeus straight. Longitudinally wrinkled near the supraorbital seta, transversally wrinkled on the forehead. Eyes prominent. One supraorbital seta, rarely asymmetrically two setae. Vertex impunctate. Antennae long and slender extending to the middle of the elytra. Antennal scape elongate, subcylindrical, basally narrowed, as long as the eye's diameter, with one dorsal seta. 2 nd antennomere with one seta ventroapically. Maxillary stipes flat, laterally with robust setae. Penultimate labial palpomere trisetose. Mentum with a bifid medial tooth. Submentum with a row of 12–14 setae. Microreticulation of the head isodiametric.

Pronotum transverse, ratio width/length of the pronotum = 1.36. Lateral margin convex, narrowed basally, widely rounded to the anterior angles, evenly and convexly narrowed to the posterior angle, faintly concave just in front of the basal angles ( Fig. 31A View Fig ). Posterior angles distinctly narrower than the anterior ones. Lateral explanation narrow, impunctate, basally opened out into the deep basal groove. Lateral margin basally blade-like, obliquely upturned. Anterior angles wide, rounded, not protruding. Posterior angles obtuse-angled. Basal margin straight, not sinuate (or only faintly sinuate). Pronotal disc convex. Basal fovea deep, anterior and posterior transverse sulci distinct, median line shallow. Basal fovea, anterior and posterior transverse sulci sparsely punctate. Apical margination of the pronotum restricted to lateral one-fourths, merges with a distinct ridge at the anterior angle; the ridge continues along the lateral groove to the hindmost lateral seta. Base of the pronotum not margined. Basolateral seta present. 2–3 midlateral setae in the apical half of the pronotum, rarely unisetose; midlateral setae inserted at the inside of the slighty upturned margin. Microreticulation of the pronotum isodiametric, impunctate on disc. Proepisternum smooth. Prosternal process triangular, flattened, margined laterally and unmargined at apex, bulging at apex.

Elytral outline moderately convex, ovoid-elongate, drop-like, narrowed basally, widest in the apical half of the elytra; lateral margin near the mesofemur straight or faintly concavely sinuate; subapically faintly sinuate. Elytral apex rounded. Basal margination straight, joined at an obtuse angle with the lateral margination. Humeral carina faintly developed. Striae distinct on disc, faintly punctate. Striae obliterate towards the apex. Intervals rather convex, more convex on disc and towards the basal margin. Interval 3 with 4–8 setae, intervals 5 and 7 with 0–3 setae. 1 scutellar seta present, rarely bilaterally with two setae. Microreticulation oblong. Mesepisterna smooth. Metepisterna smooth, twice as long as wide. Metacoxa with 2(–3) basal and one apical setae.

Sternum 2 laterally smooth. Sternum 3 medially asetose, rarely with one seta. Sterna 4–6 each with 3–6 posterior paramedial setae. Anal sternum with 1–2 paramedial setae in the male and 2–6 in the female. All sterna with faint impressions laterally.

Legs long and slender. All tarsi dorsally glabrous. Protarsus of the male with tarsomeres 1–3 distinctly broadened, ventrally with pads of adhesive setae. Ventroapical tooth of metatarsomere 4 long, longer than half of the dorsal metatarsomere length, bearing long setae. All tarsomeres 5 ventrally with two rows of 3–5 short setae. Metatarsomere 5 longer than the metatarsomeres 3+4.

Male genitalia: Edeagus ( Fig. 20A View Fig ): Basal part of the median lobe wide, with distinct basolateral membranous lobes. Mid-shaft slender, one third from the tip on the ventral side faintly bent. Tip of the apex short. Apex faintly deflected to the left. Mid-shaft of the everted endophallus densely and extensively covered with small spines.

Female genitalia: Gonocoxa ( Fig. 10A View Fig ): Large, gonocoxite 1 long. Gonocoxite 2 elongate-triangular, longer than half of length of the proximal gonocoxite, broad at base, apex broadly rounded, dorsally arcuate, dorsally distinctly grooved. Ventral preapical insertion furrow elongate-oval, with two nematiforme setae. Gonocoxites ventrally unjointed and continuously sclerotized, dorsally separated by a membranous area.

Body ratios: hea.w/fro.w = 1.43±0.023 (1.38–1.47); prm.w/hea.w = 1.26± 0.043 (1.15–1.30); prm.w/prp.w = 1.68±0.028 (1.63–1.72); pra.w/prp.w = 1.17± 0.043 (1.09–1.24); ely.w/prm.w = 1.60±0.043 (1.53–1.69); prm.w/prm.l = 1.34± 0.045 (1.24–1.40); ely.l/ely.w = 1.62±0.040 (1.55–1.70).

Etymology: The specific epithet refers to the three (the 3 rd, 5 th and 7 th) oligosetose intervals of the elytra.

Diagnosis: N. triseriata sp. nov. and N. martensi are the only Epinebriola species with the 5 th and 7 th intervals of the elytra setose. N. triseriata differs from N. martensi by the trisetose penultimate labial palpus (bisetose in N. martensi ), by the presence of a scutellar seta, which generally is absent in N. martensi , by the distinct striae of the elytra (shallow in N. martensi ), and by the mid-shaft of the endophallus densely covered with setae (asetose in N. martensi ).

Distribution ( Fig. 41 View Fig ): Known only from the type locality on western slope of the Annapurna South Himal, western Central Nepal Himalaya.

Habitat: All specimens were collected alongside a melt water stream in the high alpine zone.