Eratomyia magnifica, Published, 2007

DE SOUZA AMORIM, DALTON & RINDAL, EIRIK, 2007, Phylogeny of the Mycetophiliformia, with proposal of the subfamilies Heterotrichinae, Ohakuneinae, and Chiletrichinae for the Rangomaramidae (Diptera, Bibionomorpha), Zootaxa 1535 (1), pp. 1-92 : 14-15

Eratomyia magnifica , sp. n.

Diagnosis. As for the genus.

Material examined. Holotype, ♂, ECUADOR, Quito-Baiza, E. Papallactam, 2,900m, 12–15 II 1971, L.G Peña col. ( MZSP).

Description. Total length, 5.58 mm. Wing length, 4.80 mm. Terminalia length, 0.22 mm.

Male. Head ( Fig. 7 ). Scape yellow. Antenna dark brown, 1st flagellomere 1.5 times longer than 2nd flagellomere; 4th flagellomere 4.4 times longer than wide, densely setose (both flagella broken). Frons light brown with some setae. Palpi yellow, except darker segment 1, very long, twice length of head, palpiger plus 4 palpomeres, last segment 1.5 longer than penultimate. Labella yellow. Stipes long, shining yellow. Occiput dark brown. Thorax ( Fig. 8 ). Scutum shining blackish brown, with pair of rows of dorsocentral setae. Scutellum blackish brown. Pleural sclerites shining blackish brown, except for katepisternum, covered with silvery pruinescence, and brownish episternum II. Halter brownish yellow, base longer than scape. Episternum I not particularly developed. Epimeron I triangular. Anepisternum bare, anepisternal cleft deep, suture reaching primary anapleural suture. Katepisternum with pair of setae dorsally. Suture between katepisternum and laterotergite nearly vertical. Midpleural pit absent. Mesepimeron long, with 9 setae, suture extending from dorsal margin. Suture between laterotergite and mediotergite deeply concave. Mediotergite high, only slightly curved. Metepisternum well developed, metepimeron discernible. Mesopleurotrochantin apparently absent, a small, triangular sclerite present at contact between katepisternum, mesepimeron, and anterodorsal extension apparently not homologous. Legs. Coxae long, brownish yellow at base, trochanters brownish, femora yellow, long, tibiae ocher-brown, tarsi brown. Spurs 1-2-2. Tibia 1 with spur on inner ventral apex, and modified area at inner apex bearing regular row of setae ( Fig. 9 ). Meron clearly present on coxa II, in format similar to that in some limoniids and in pachyneurids. Wing ( Fig. 10 ). Sc incomplete, reaching basal third of wing. R 1 long, reaching C beyond distal third. R 5 reaching C close to wing apex, running quite apart from R 1. r-m long, slightly longer than first sector of Rs. Medial fork short, M 1+2 separating from M 3+4 beyond half of wing. Basal cell wide, m-cu and M 1+2 / M 3+4 quite separated from base of R; m-cu long, gently curved. A 1 incomplete, absent on distal third, anal lobe reduced. Abdomen. Tergites 1–4 and 6 dark shining brown, tergite 5 yellow. Tergites wide, sternite narrow. Terminalia ( Figs 11–15 ). Terminalia black. Segment 8 normally developed. Sternite 9 triangular, laterally fused to gonocoxite to form synsternogonocoxite. Gonocoxites not projected beyond gonostyles, pair of digitiform processes arising inward distally, bearing some setae and pair of spines. Gonostyles more or less digitiform, directed inward, setose (without spines), with short tooth at apex. Aedeagus present with apodeme proximally and tegmen-like structure distally. Gonocoxal bridge well developed dorsal to aedeagus. Tergite 9 with pair of well-developed distal projections and deep mesal incision almost reaching anterior margin; projections with about 16 black spines. Tergite 10 present, weakly sclerotized, apparently fused to cerci (or with ear-like distal projections laterally).

Female. Unknown.

Etymology. The species name comes from the Latin word magnificus, meaning magnificent or splendid.

Figures

FIGURES 7–10. Eratomyia magnifica, sp. n. 7. Head (apical maxillary palpal segments not represented), lateral view. 8. Thorax. 9. Front tibial apex, inner view. 10. Wing.

FIGURES 11–15. Eratomyia magnifica, sp. n. 11. Male terminalia, posterior view. 12. Male terminalia, lateral view. 13. Gonostyle, external view. 14. Aedeagus and gonocoxal bridge. 15. Tergites VIII–X and cerci, posterior view.