Patrera procera ( Keyserling, 1891 )

Oliveira, Luiz Fernando M. & Brescovit, Antonio D., 2021, The spider genus Patrera Simon (Araneae: Dionycha, Anyphaeninae) in the Atlantic Forest biome, with the description of one new species from Brazil, Zootaxa 4991 (2), pp. 201-246 : 203-210

Patrera procera ( Keyserling, 1891)

Figs 1A–G ; 2A–D ; 3A–F ; 27C

Anyphaena procera Keyserling, 1891: 86 , figs 49–49a (Syntypes male and female from Rio de Janeiro (22°54’10”S; 43°12’28”W),

Rio de Janeiro, Brazil, 7.I.1890, Keyserling leg., deposited in BMNH 1890.7.1.613-614, examined. Lectotype male and paralectotype female here designated). Teudis procerus: Petrunkevitch, 1911: 518 . Aysha cinereovittata Mello-Leitão, 1945: 242 (partim, only the females MLP; males = Iguarima censoria (Keyserling) , see

Brescovit, 1997: 50. Patrera procera: Brescovit, 1997: 33 , 50.

Other material examined. BRAZIL. No date, 1 ♂ (MNHN 18666) ; Paraíba: João Pessoa, Mata do Buraquinho (7°8′17″S; 34°50′42″W), 1♀, 2003–2004, S.C. Dias leg. (IBSP 99968) ; Areia, Reserva da Mata do Pau Ferro (06º57’S; 35º44’W), 1♂, 23–29.IX.1999, A.D. Brescovit et al. leg. (IBSP 138929); 2 ♂ (IBSP 138939); 1♂ (IBSP 138949); 1♂ (IBSP 138950); 1♂ (IBSP 138958); 1♂ (IBSP 138960); all collected in 23–29.IX.1999 by A.D. Bresco- vit et al .; Bahia: 2♂ (MNHN 7135); Una, Reserva Biológica do Una (15°10′23″S; 39°7′56″W), 1♀, 15–28.XI.2000, A.D. Brescovit et al. leg. (IBSP 45700); Jussari (15°11’27”S; 39°29’42”W), 3♀, 1970, Cepec / Ceplac leg. (MNRJ 3143) ; 1♂, 1♀ (MNRJ 3236); 1♂ (MNRJ 3194); 2♂ (MNRJ 3256); 1♂, 2♀ (MNRJ 13313); 2♂ (MNRJ 3183); Coaraci (14°38′27″S; 39°33′3″W), Fazenda Boa Esperança, 1♂, 01.X.1970, Cepec/Ceplac leg. (MNRJ 3231); Mato Grosso do Sul: Bonito , Baia Bonita (21º09’S; 56º26’W), 1♂ (IBSP 138887); 1♀ (IBSP 138890); 3♂, 2♀ (IBSP 138902); all collected in 14–23.X.2002, Equipe Biota ; Minas Gerais: Belo Horizonte, Estação Ecológica UFMG, Malaise trap (19°52′19″S 43°57′58″W), 2♂, 09–16.XI.1999, J.C.R. Fontenelle leg. (IBSP 41119) ; Espírito Santo: São José do Calçado, Rio Itabapoana, Usina Hidrelétrica de Rosal (20°57’7”S; 41°43’5”W), 1♂, 1♀, XI.1999, I. Knysak leg. (IBSP 26383) ; Apiacá (21°9′14″S; 41°34′4″W), 1♂, XI.1986, R.L.C. Baptista leg. (MNRJ 3188) ; Ca- riacica, Alto do Alegre, Reserva Estadual de Duas Caras (20°17’29.7”S; 40°31’10.2”W), 1♂, 13.X.2005 , T. Souza et al. leg. (IBSP 133182); Linhares, Flona de Goytacazes (19°23’48”S; 40°3’42”W), 42m alt., 2♂, 21–23.X.2005 , T. Souza et al. leg. (IBSP 134825); Conceição da Barra, Floresta Nacional do Rio Preto , Trilha do Mata-Pau , 60m alt. (18°24’00”S; 39°50’00”W), 1♂, 19.X.2005 , T. Souza et al. leg. (IBSP 135120); Rio de Janeiro: Volta Redonda, Floresta da Cicuta (22°32′53″S; 44°5′34″W), 3♀ , VIII.2002, E.L.C. Silva et al. leg. (IBSP 48942; IBSP 48950; IBSP 48954); Cachoeiras de Macacu, Reserva Ecológica de Guapiaçu (22°35′14.61″S; 42°56′24.69″W), 11♂ (IBSP 138968; IBSP 138970; IBSP 138972; IBSP 138974; IBSP 138984; IBSP 138991; IBSP 138993; IBSP 138994; IBSP 138997); all collected in 08–12.X.2001 by Equipe Biota ; Pinheiral , Fazenda Santa Helena (22º34’00”S; 44º21’00”W), 21♂, 3♀ (IBSP 139401; IBSP 139406-IBSP 139408; IBSP 139410; IBSP 139411; IBSP 139414; IBSP 139415; IBSP 139418-IBSP 139420; IBSP 139423-IBSP 139427; IBSP 139430; IBSP 139432- IBSP 139434); all collected in 05–11.X.1999 by A.D. Brescovit et al.; Pinheiro (currently Piraí , Pinheiral ) (22°30′46″S; 44°00′3″W), 2♂, I.1988, R.L.C. Baptista leg. (MNRJ 3189) ; 1♂, 1♀, 4#imm. [without date], C.F. de Mello-Leitão leg. (MNRJ 616); São Paulo: Teodoro Sampaio, Porto Cabral, Rio Paraná (24°31’00.1”S; 47°42’00”W), 1♀, 1941, L.P. Travassos Filho leg. (MZSP 9788) ; Ribeirão Preto (21°10′40″S; 47°48′36″W), 1♂, 1992, E. Camillo leg. (IBSP 210237) ; Itapetin- inga, Estação Ecológica de Angatuba (23º24’00”S; 48º21’00”W), 2♂, 7♀ (IBSP 138710; IBSP 138711; IBSP 138714; IBSP 138717; IBSP 138720; IBSP 138722; IBSP 138723; IBSP 138725), all collected in 11–16.XI.2002 by Equipe Biota; Iporanga (24°35’09”S; 48°35’34”W), Lajeado, 1♂ , XII.1944, J. Damico leg. (MZSP 12217); Itaberá (23°51′43″S; 49°8′13″W), 2♂, 1♀, XI.1943, F.S. Pereira leg. (MZSP 12207) ; Botucatu, Fazenda Butignoli, Rubião Júnior (23°59’21”S; 48°26’21”W), 1♀, 07.I.1987, I.M.P. Rinaldi & L.C. Forti leg. (MCN 22486) ; 1♂, 07.X.1987, I.M.P. Rinaldi & L.C. Forti leg. (MCN 22485) ; 1♀, 12.II.1987, I.M.P. Rinaldi leg. (MZSP 60386); 2♀, 18.XII.1978, N. Carneiro leg. (MZSP 12129); 1♀, 02.XII.1987, I.M.P. Rinaldi & L.C. Forti leg. (MCN 22483) ; 1♂, 02.XII.1987, I.M.P. Rinaldi & L.C. Forti leg. (MCN 22484) ; 3♂, 1♀, 14–16.XII.1966, (MZSP 12123; MZSP 12141; MZSP 12317; MZSP 14137); all collected by V.C. de Jesus; 1♂, 16.XII.1986, I. Rinaldi leg. (MZSP 60360); Botucatu (22°53’9”S; 48°26’42”W), Parque Municipal , 1♀, 11.V.1986 (MZSP 60383); 1♂, 12.II.1987 (MZSP 60385); 1♂, 13.I.1988 (MZSP 60361); 3♀, 02.VIII.1988 (MZSP 60362); 3♀, 12.II.1987 (MZSP 60363), all collected by I. Rin- aldi; Fazenda Edgárdia, campus UNESP (22°48’56”S; 48°23’44”W), 1♀, 07.VI.1988, I.M.P; Rinaldi leg. (MZSP 60384) ; Rio Claro (22°24′39″S; 47°33′39″W), 1♀, 17–19.I.1962, H.R. Filho leg. (MZSP 10216) ; Assis, Estação Ecológica de Assis (22º34’00”S; 50º24’00”W), 1♂, 3♀, 25–30.XI.2002, Equipe Biota leg. (IBSP 138827; IBSP 138842; IBSP 138860) ; Atibaia (23º07’01’’S; 46º33’01”W), 1♂, 13.X.2002, G. Machado et al. leg. (IBSP 35177); 1♂, X.2002, M.B. Rosseto & G. Machado leg. (IBSP 35845) ; Biritiba Mirim (23°34′22″S; 46°2′20″W), 1♂, 2003, I. Knysak leg. (IBSP 72718) ; Jundiaí (23º11’11”S; 46º53’03”W), 1♂, 09–12.X.1999, C.A. Rheims leg. (IBSP 26252) ; Reserva Biológica Municipal da Serra do Japi (23°12’00”S; 46º30’00”W), 2♂, 2004, Alunos Curso Pós USP leg. (IBSP 138871; IBSP 138872); 1♀ , III.2007, A.D. Brescovit et al. leg. (IBSP 215510); 1♂, 2♀, 15– 17.XII.2014, Exc. Alunos USP leg. (IBSP 231027); Cabreúva (23º18’27”S; 47º07’58’’W), 27.X.2001, D.F. Candi- ani & P.E.D. Francesconi leg. (IBSP 31976) ; Itapecerica da Serra (23°43′1″S; 46°50′56″W), 1♂, 10.I.2006, L.V. de Camões leg. (IBSP 75579) ; Mogi das Cruzes, Rio Tietê (23°31′22″S; 46°11′16″W), 1♀, VII–VIII.1996, R. Martins leg. (IBSP 13831) ; Barueri (23°30′38″S; 46°54′34″W), 1♀, 07.I.1961 (MZSP 12250); 2♀, 31.XII.1965 (MZSP 14108); 2♀, 21.XII.1965 (MZSP 14139), 1♂, 08.I.1966 (MZSP 5958); 1♂, 08.I.1966 (MZSP 12314); 3♂, 1♀, 31.XI.1965 (MZSP 14136); 1♂, 1♀, 05.II.1965 (MZSP 14138); all collected by K. Lenko ; Embu das Artes (23°38′56″S; 46°51′7″W), 1♀, II.1946, F. Lane leg. (MZSP 12242) ; São Paulo (23º32’51”S; 46º38’10”W), 1♀, X.1990, Equipe IBSP leg. (IBSP 4367); Vila Mascote (23°38’44”S; 46°40’2”W), 1♀, 04.XI.2002, D.M.B. Battesti leg. (IBSP 36392) ; Perdizes (19°21’19’’S; 47°16’58’’W), 1♂, 01.XII.1951, H. Urban leg. (MZSP 12276) ; Parque da Previdência (23º34’00”S; 45º43’00”W), 1♂, 22.X.1999, F.S. Cunha leg. (IBSP 29400) ; 1♂, 29.X.1999, J. Im- pério leg. (IBSP 29401); Campus USP, Reserva Florestal da CUASO (23º33’00”S; 46º43’00”W), 3♀, 14.XII.1999 (IBSP 29393); 1♀, 29.X.1999 (IBSP 29394); 1♂, 29.X.1999 (IBSP 29395); 1♀, 31.X.2000 (IBSP 29396); 1♀, 14.XII.1999 (IBSP 29398); 1♂, 27/X/1999 (IBSP 32984); 2♀, all collected by F.S. Cunha ; 29.X.1999, D.F. Candi- ani leg. (IBSP 29399); 1♂, 05.IX.2000, J. Império leg. (IBSP 29397); Campus Instituto Butantan (23º33’00”S; 46º43’00”W), 4♂, 2♀, 11.XI.2004, no coll. (IBSP 48427); 2♀, 2010, G.S. Ruiz leg. (IBSP 168580) ; Campus Insti- tuto Butantan, Mata IBSP, 5♂, 6♀, 05.X.2005, Alunos USP curso 2006 leg. (IBSP 57444); Represa de Guarapir- anga, Ilha dos Eucaliptos (23°44’26”S; 46°44’09”W), 1♂, 09.IX.2001, R.P. Indicatti leg. (IBSP 32580) ; Ilha dos Eucaliptos, 4♂ 8♀ (IBSP 61264; IBSP 61265; IBSP 61266; IBSP 61267; IBSP 61268; IBSP 61269; IBSP 61270; IBSP 61271; IBSP 61272); all collected in 07–13.X.2004 by I. Cizauskas & C.R.M Garcia; Parque Estadual do Jaraguá (23°27′34.3″S; 46°46′2.8″W), 2♂, 2♀, 27–28.X.2005, Curso UFRRJ leg. (IBSP 71185; IBSP 71237; IBSP 71239); 10♂, 4♀, 14–19.X.2002, Equipe Biota leg. (IBSP 139596; IBSP 139598; IBSP 139599; IBSP 139602; IBSP 139603; IBSP 139605; IBSP 139615; IBSP 139616; IBSP 139618; IBSP 139624; IBSP 139636; IBSP 139647; IBSP 139648) ; Parque Estadual da Cantareira, Sede (23°24′5″S; 46°35′24″W), 1♀, 20.XII.2000 (IBSP 85701); 2♀, 21.XII.2000 (IBSP 85702); 1♀, 28.II.2001 (IBSP 85703); 2♂, 15.XII.2000 (MCN 41880; MCN 41882); 2♂, 3♀ (MZSP 24966–MZSP 24970), all collected in 22.XII.2000 by R. Pinto da Rocha et al.; Núcleo Pedra Grande, 1♀, 15.XII.2002, R. Pinto da Rocha et al. leg. (IBSP 210221); 3♂, 3♀ (MZSP 24959–MZSP 24964), all collected in 21.XII.2000 - 6.I.2001 by R. Pinto da Rocha et al.; Parque Estadual Fontes do Ipiranga (23°38′24.6″S; 46°37′3.1″W), 1♀, 28.X.2017 (IBSP 222758); 1♂, 27.VII.2018 (IBSP 222766); 1♀, 28.X.2017 (IBSP 222768); 1♀, 25.XI.2017 (IBSP 222769); 1♂, 30.IX.2017 (IBSP 222777); 1♂, 27.VIII.2017 (IBSP 222778); 1♂, 22.VII.2017 (IBSP 222780); 1♀, 25.XI.2017 (IBSP 222783); 1♂, 30.IX.2017 (IBSP 222787); 1♂, 27.VIII.2017 (IBSP 222792); 1♀, 27.VIII.2017 (IBSP 222793); 1♀, 27.VIII.2017 (IBSP 222794), all collected by Z. R. Mendes ; Horto Florestal (currently Parque Estadual Alberto Löfgren ) (23°27′33″S; 46°38′2″W), 1♂, 1imm., 1.XI.1994, R. Pinto da Rocha leg. (MZSP 14310) ; Bosque da Saúde (23°34’21”S; 46°38’70”W), 2♀, 09.IV.1944, F. Lane leg. (MZSP 5165) ; 1♂, 22.III.1942, F. Lane leg. (MZSP 14134) ; Cocaia, Represa Billings (23°47’13”S; 46°35’2”W), 1♀, 11.III.1951, H. Urban leg. (MZSP 12212) ; Parque Estadual Serra da Cantareira, Sede (23°24′5″S; 46°35′24″W), 1♂, 22.XI.2000, R. Pinto da Rocha et al. leg. (MZSP 25011); São José do Barreiro , Parque Nacional da Serra da Bocaina (23°2′30″S; 44°39′42″W), 1♂, 22–25.VII.2006, F.U. Yamamoto leg. (IBSP 66747) ; Ribeirão Grande, Parque Estadual de Intervales (24°16′6.3″S; 48°24′49.7″W), in Bamboo, 1♀, 2002–2003, M.O. Gonzaga leg. (IBSP 71452) ; São Bernardo do Campo (23°41′38″S; 46°33′54″W), Clube de Campo São Camilo , 1♂, 1♀, 20.X.2008, V. Passanha & C.V. Janini leg. (IBSP 119790) ; Mairiporã (23°22′44″S; 46°31′38″W), Serra da Cantareira , 1♂, VII.2000, C.R. Macagnan leg. (IBSP 27347) ; Mogi Guaçu, Reserva Biológica Mogi Guaçu (22º10’00”S; 47º08’00”W), 2♀, 16.XII.1998, E. Camillo leg. (IBSP 28113; IBSP 28155) ; Reserva Biológica Mogi Guaçu (22º10’00”S; 47º08’00”W), 33 ♂, 2 ♀ (IBSP 54646; IBSP 139460; IBSP 139466; IBSP 139468; IBSP 139469; IBSP 139473; IBSP 139482; IBSP 139484; IBSP 139486; IBSP 139489; IBSP 139491; IBSP 139493; IBSP 139497; IBSP 139500; IBSP 139501; IBSP 139502; IBSP 139504; IBSP 139505; IBSP 139509; IBSP 139510; IBSP139511; IBSP 139512; IBSP 139514; IBSP 139516; IBSP 139517; IBSP 139518), all collected in 13–19/X/2003 by Equipe Biota; Pariquera-Açu (24°42’54”S; 47°52’51”W), Vale do Ribeira , cam- pus do Instituto Agronômico de Campinas , 1♂, [without date], E. Wienskoski leg. (MZSP 60391) ; Ubatuba (23º26’02”S; 45º04’16”W), Fazenda Experimental , 1♂, 1♀, 14.XI.1943, A. Zoppei leg. (MZSP 12228) ; Paraná: Cruzeiro do Iguaçu, Usina Foz do Chopim (25º33’45”S; 53º10’37”W), 1♂, 08–15.X.1998, Equipe IBSP leg. (IBSP 21241); Adrianópolis , Vale do Ribeira (24°39′25″S; 48°59′27″W), 1♀, 18.XI.2004, D.M.B. Battesti leg. (IBSP 47949) ; Bela Vista do Paraíso (22º59’49”S; 51º11’27”W), 1♂, 1♀, 11–16.XI.1991, A.B. Bonaldo leg. (IBSP 221089) ; Fênix (23º54’57”S; 51º58’44”W), 1♂, 24.XI.1986, Equipe CIIF leg. (IBSP 221097); 1♂, 01.XI.1986, Equipe Profaupar leg. (IBSP 221098) ; Parque Estadual de Vila Rica do Espírito Santo (23°55’00”S; 51°57’00”W), 3♀, 12.XI.1987, A.D. Brescovit leg. (IBSP 221096) ; Curitiba, campus UFPR, Centro Politécnico (25°25′44″S; 49°16′3″W), 2♂, 2♀, 26.X.1995, A.B. Bonaldo leg. (MCN 26698) ; Almirante Tamandaré (25°19′30″S; 49°18′36″W), 1♀, 10.VII.1984 (MCN 12389); 1♂, 25.IV.1984 (MCN 12414); 1♂, 08.VI.1984 (MCN 12454); 1♂, 25.IV.1984 (MCN 12467); 1♂, 22.VI.1984 (MCN 12488); 1♂, 28.IX.1984 (MCN 12517); 1♂, 28.IX.1984 (MCN 12518); all collected by E. C. Costa ; Capanema (25°40′19″S; 53°48′32″W), 3♂, 11.III.1981, S. Scherer leg. (MCN 9777) ; Santa Catarina: Chapecó, Floresta Nacional de Chapecó (27º06’10,5”S; 52º46’48,8”W), 1♀, I.2003 – XII.2004, M. Scartezini leg. (IBSP 91070) ; Nova Teutônia (27°11’00”S; 52°23’00”W), 2♂, 1♀, [without date], F. Plaumann leg. (FMS 13579; FMS 13583; FMS 13578) ; Florianópolis, Itacorubí , UCAD (27°33′21.44″S; 48°29′37.46″W), 1♂, 2004, J. Steiner leg. (IBSP 91097) ; Criciúma (28°40′40″S; 49°22′12″W), 1♂, 08.XI.2007, R.A. Teixeira leg. (IBSP 132769) ; Rancho Queimado (27°40′22″S; 49°1′19″W), 1♂, 15–18.XI.1995, L.A. Moura leg. (MCN 27039) ; 2♀, 13–15.I.1995, A.B. Bonaldo leg. (MCN 26356) ; Rio Grande do Sul: Iraí (27°11′38″S; 53°15′3″W), 1♂, 2♀, 20.XI.1975, A.A. Lise leg. (MCN 3106) ; Derrubadas, Parque Estadual do Turvo, Salto do Yucumã (27°08’32.5″S; 53°53’09.0″W), 1♀, 16.I.1985 (MCN 13002); 1♂, 16.I.1985 (MCN 13005); all collected by A.A. Lise ; São Fran- cisco de Paula (29°26′52″S; 50°35′2″W), 1♀, 22.IX.1974 (MCN 2304); 1♂, 1♀, 22.I.1981 (MCN 9531); 1♂, 2♀, 23.I.1981 (MCN 9540); all collected by A.A. Lise ; Barragem dos Bugres (29°20’35.87”S; 50°41’44.15”W), 2♂, 23–25.XI.1998, A.F. Franceschini leg. (MCN 30656) ; 2♂, 1♀, 01–04.II.1999, L.A. Moura leg. (MCN 30669) ; Rincão dos Kroeff (29°28’3”S; 50°20’47”W), 1♀, 05.I.1985, A.A. Lise leg. (MCN 12739) ; Cambará do Sul, Itaim- bézinho (29º02’52”S; 50º08’41”W), 2♂, 1♀, 06.I.1985, A. Lise leg. (MCN 12777; MCN 12779; MCN 12780) ; Garabi (28°12’50”S; 55°41’17”W), 1♀, XI.1988, A.A. Lise leg. (IBSP 258334) ; Marcelino Ramos, Estreito Au- gusto Cesar (27°24’51”S, 51°27’2”W), 1♀, 03.II.1990, C.A. Martinazzo leg. (MCN 19530) ; Maquiné (29°40’30”S; 50°12’25”W), Fepagro, 1♀, 18–27.I.2002, Equipe Biota leg. (IBSP 139226) ; Derrubadas, Parque Estadual do Turvo (27°8’44”S; 53°53’10”W), 6♀ (IBSP 139987; IBSP 139989; IBSP 139990; IBSP 140007; IBSP 140018), all col- lected in 11–18.I.2002 by Equipe Biota ; Parque Estadual do Turvo (27°8’44”S, 53°53’10”W), 12♂, 3♀, 27– 31.X.2002 (MCN 37768); 1♂, 1♀, 19–22.X.2004 (MCN 38812); 1♂, 27–31.X.2003 (MCN 38813); 6♂, 27–31. X.2003 (MCN 38814); 1♂, 19–22.X.2004 (MCN 38836); 10♂, 6♀, 19–22.X.2004 (MCN 38841), all collected by R. Ott et al.; Canela (29°21’57”S; 50°48’57”W), 1♀, 31.XII.1973 (MCN 2070); 1♂, 20.III.1976 (MCN 3854); 1♀, 05.III.1977 (MCN 5209); 1♀, X.1976 (MCN 6300), all collected by A.A. Lise ; 1♂, 1♀, 03–04.IX.1988, A.B. Bon- aldo leg. (MCN 17841); 1♀, 05.II.1973 (MCN 852); 2♂, 31.XII.1973 (MCN 2044); 1♀, 11.V.1974 (MCN 2163); 1♂, 26.XII.1974 (MCN 2475); 1♂, 1♀, 26.II.1975 (MCN 3748); all collected by A.A. Lise ; Carazinho (28°17’2”S; 52°47’9”W), 2♂, 10.XI.1979, H. Bischoff leg. (MCN 8655) ; 1♀, 10.XI.1979, A.A. Lise leg. (MCN 8686) ; So- bradinho (29°25’15”S; 53°1’44”W), 2♀, 10.I.1985, A.A. Lise leg. (MCN 18717) ; Marcelino Ramos, Estreito Au- gusto Cesar (27°24’51”S; 51°27’2”W), 1♀, 03.II.1990, C.A. Martinazzo leg. (MCN 19529) ; Nova Petrópolis, Boê- mios, Linha Imperial (29°21’36”S; 51°2’54.8”W), 1♂, 20.XII.1973, A.A. Lise leg. (MCN 2206) ; Caxias do Sul, Vila Oliva (29°12’56”S; 50°53’22”W), 1♂, 10.I.1976, P.A. Buckup leg. (MCN 3959) ; Caxias do Sul (29°10’04”S; 51°10’44”W), Água Azul, 1♀, 15.IX.1976, H. Bischoff leg. (MCN 4532) ; 1♂, 04.IV.1975, A.A. Lise leg. (MCN 18708) ; General Câmara (29°54’18”S; 51°45’36”W), 1♂, 1♀, 14.X.1982, C.J. Becker leg. (MCN 10810) ; Far- roupilha (29°13’30”S; 51°20’52”W), 2♂, 1♀, 29.IX.1978, H. Bischoff leg. (MCN 8283; MCN 8306; MCN 8329); Agudo, Usina Hidrelétrica Dona Francisca (29°27’7”S; 53°16’55”W), 1♀, 02.XI.2002, A.B.B. Morais leg. (IBSP 122191) ; 1♀, Passo Fundo (28°15’46”S 52°24’25”W), 1♀, 13.X.1985, A.A. Lise leg. (MCN 14321) ; 1♂, 01.VIII.1986, A.A. Lise leg. (MCN 15573) ; Estrela Velha, Barragem de Itaúba (29°15’40”S; 53°14’8”W), 2♂, 1♀, 21.X.1998, A.B. Bonaldo leg. (MCN 30606; MCN 30607) ; Garruchos (28°11’2”S; 55°38’20”W), 4♀, 10.XII.1975 (MCN 3152); 3♀, 10.XII.1975 (MCN 3172); 1♀, 06.XII.1975 (MCN 18703); all collected by A.A. Lise ; Triunfo (29°56’34”S; 51°43’4”W), 2♂, 1♀, 15.IX.1977, H. Bischoff leg. (MCN 6516; MCN 6969) ; 1♂, 27.X.1977, M.H. Galileo leg. (MCN 7031) ; 1♂, 2♀, 28.XI.1977, A.A. Lise leg. (MCN 7393); Polo Petroquímico do Sul (29°54’4”S; 51°22’27”W), 1♂, 1♀, 20.X.1977, E.H. Buckup leg. (MCN 18706; MCN 18707) ; Parobé (29°37’44”S; 50°50’6”W), Rio dos Sinos, 1♀, 10.XI.2008, E.N.L. & P.E.S. Rodrigues leg. (MCN 47891) ; 1♀, 18.XII.2007, E.N.L. & P.E.S. Rodrigues leg. (MCN 48029) ; Parque Braskem de Proteção Ambiental (29°51’59.44”S; 51°21’54.00”W), 2♂, 24.X.1989, A.D. Brescovit leg. (MCN 18903) ; 1♂, 28.XI.1989, E.H. Buckup leg. (MCN 19042) ; 1♀, 24.XI.1999, M.A.L. Marques leg. (MCN 31739) ; 1♂, 21.X.2003, A. Barcellos & L. Schmidt leg. (MCN 36458) ; 1♀, 2.XI.2007, A. Barcellos & L. Schmidt leg. (MCN 43644) ; 1♀, 04.XII.2009, M.R.M. Poeta leg. (MCN 46917) ; 2♂, 04.XII.2009, M. Pairet Jr, leg. (MCN 46925); 1♀, 04.XII.2009, P.E.S. Rodrigues leg. (MCN 46933) ; Arroio Grande, Mata Ciliar do Rio Piratini (31°54’06.47”S; 52°39’08.29”W), 1♀, 13.VI.2008, E.N.L. & P.E.S. Rodrigues leg. (MCN 54732) ; Campo Bom (29°40’44″S; 51°3’10”W), 1♀, 16.XI.1979, C.J. Becker leg. (MCN 8789) ; Novo Hamburgo (29º40’40”S; 51º07’51”W), 2♂, 22.IX.1986, C.J. Becker leg. (IBSP 221094) ; 1♂, 21.X.1983 (MCN 11814); 1♀, 25.XI.1983 (MCN 11847); 2♂, 11.XII.1984 (MCN 12645); 2♂, 07.X.1985 (MCN 14262; MCN 12646); 2♂, 14.XII.1984 (MCN 12717); 1♀, 31.III.1986, C.J. Becker leg. (MCN 14557) , all collected by C.J. Becker; Morro Reuter (29°32’16”S; 51°4’51”W), 1♀, 28.III.1976, C.J. Becker leg. (MCN 3576) ; Viamão, Estação Experimental Fitotécnica Águas Belas (30°05’00”S; 51°02’00”W), 1♂, 30.XI.1984, M.A.L. Marques leg. (MCN 12611) ; 1♂, 06.I.1977, A.A. Lise leg. (MCN 5899); São Leopoldo (29º45’36”S; 51º08’49”W), 1♂, 28.IX.1987, C.J. Becker leg. (IBSP 221095) ; 2♂, 17.IX.1982 (MCN 10965); 3♂, 24.IX.1982 (MCN 11049); 2♂, 15.X.1982 (MCN 11146); 1♀, 15.X.1982 (MCN 11147); 1♀, 20.XII.1982 (MCN 11339); 2♂, 12.XII.1983 (MCN 11865); 1♂, 26.XII.1983 (MCN 11911); all collected by C.J. Becker ; Montenegro (29°41’20”S; 51°27’39”W), 2♂, 2♀, 06.X.1977 (MCN 6806; MCN 7277); 3♂, 17.XI.1977 (MCN 7278); 1♀, 20.XII.1985 (MCN 14077); 4♂, 29.IX.1977 (MCN 6718); 1♂, 17.XI.1977, (MCN 7280); 8♂, 01.XII.1977 (MCN 7482); 4♂, 20.XII.1977 (MCN 7667); all collected by E.H. Buckup & A.A. Lise ; Tapes (30°40’22”S; 51°23’45”W), Fazenda Guará, 1♀, 19.XII.2003, Equipe Probio leg. (MCN 36821) ; Fazenda São Miguel , 2♂, 1♀, 17.XII.2003, Equipe Probio leg. (MCN 36847) ; Barra do Ribeiro (30°17’27”S; 51°18’3”W), Fazenda Boa Vista, 2♂, 18.XII.2003, Equipe Probio leg. (MCN 36850) ; Capão do Leão, Campus UFPEL (31°45’46”S; 52°29’2”W), 1♀, 24.II.2000 (IBSP 36534); 2♂, 1♀, 30.IX.2001 (IBSP 36536); 2♂, 30.IX.2001 (IBSP 36537); 1♀, 30.IX.2001 (IBSP 36538); all collected by E.N.L. Rodrigues. PARAGUAY. Departamento Paraguari: Parque Nacional de Ibycuí (26º05’49.08”S; 56º50’20.16”W), 1♀, 28.I.1982 (MNHNP); 1♀, 20.X.1981 (MNHNP) ; Departamento Amambay: Parque Nacional Cerro Corá, Base de Cerro Tuya Hog (22°38’28”S; 56°0’47”W), 1♀, X.1983, (MNHNP); 1♀, 8.VI.1982 (IBNP) ; Departamento Itapúa: Pirapó (28°2’42”S; 55°11’56”W), 1♂, 4.VIII.1982 (MNHNP) ; Departamento Caaguazú: Caaguazú (25°28’11”S; 56°0’55”W), 1♂, 23.VII.1982 (MNHNP); all collected by J.A. Kochalka. ARGENTINA. Misiones: Loreto (27°19’55.12”S; 55°31’23.47”W), 1♂, 27.III.1932 – 8/III/1933, A.A. Oglobin leg. (MLP) ; Puerto Iguazú (25°35’50”S; 54°34’41”W), 8♀, XI.1948, M. Birabén leg. (MLP) ; Puerto Victoria (26°20’04”S; 54°39’44”W), 1♀, [without date], A. Zenzes leg. (MLP) ; Parque Nacional del Iguazú, Cataratas del Iguazú (25°41’43”S; 54°26’12”W), 2♂, XI.1989, (MACN- Ar ); 1♀, XI.1991, (MACN-Ar); 1♂, 1♀, XI.1992, (MACN-Ar); all collected by M.J. Ramírez ; Cantera , 200 m, 1♀, 8.XII.1990 – 06.I.1991, S. & J. Peck leg. (AMNH) ; Porto Canoas , 180m, 1♀, 8.XII.1990 – 06.I.1992, S. & J. Peck leg. (AMNH) ; Eldorado (26°28’00”S; 54°43’00”W), 2♂, 01.X–15.XI.1964, A. Kovacs leg. (AMNH) ; 2♂, 1♀, 01.X– 15.XI.1965, A. Kovacs leg. (AMNH).

Diagnosis. Males of Patrera procera can be distinguished from the remaining species by the characteristic tibial retrolateral apophysis wide, subrectangular, with a transverse fold at the tip, forming a conspicuous groove,

and by the conical dorsal tibial apophysis ( Figs 1D , 2B , 3E ) in the male palp. Females are diagnosed by their short, narrow epigynal lateral borders with lateral excavation at the internal borders and wide deep atrium in the epigynum ( Figs 2C , 3F ).

Description. Male (Lectotype, BMNH). Carapace orange brown, with orange paramedian stripes. Chelicerae brown yellowish, labium and endites orange. Sternum light orange, with dark borders. Legs orange. Abdomen dorsally brownish, ventrally cream ( Fig. 1A ). Total length 8.6, carapace length 3.5, width 2.65. Clypeus height 0.15. Eye diameters and interdistances: AME 0.11, ALE 0.20, PME 0.17, PLE 0.20; AME–AME 0.06, AME–ALE 0.08, PME–PME 0.21, PME–PLE 0.17, ALE–PLE 0.07. Chelicerae 2.3 long, four promarginal teeth, nine small retromarginal denticles ( Fig. 3A ). Leg measurements: leg I—femur 5.5/ patella 1.5/ tibia 6.1/ metatarsus 5.2/ tarsus 2.05/ total 20.35; II—4.85/ 1.3/ 5.25/ 4.6/ 1.9/ 17.9; III—3.1/ 1.05/ 2.5/ 3.4/ 1.0/ 11.05; IV—4.8/ 1.3/ 4.35/ 5.75/ 1.2/ 17.75. Leg spination: I—II—tibia v2-2-0, p1-1-1, r1-1-1, metatarsus v2-0-0, p1-1-0, r1-1-0; III—IV—tibia v2- 2-2, p1-1-1, r1-1-1, metatarsus v2-2-2, p1-1-1, r1-1-1. Abdomen: length 5.35, epigastric furrow 1.4 from tracheal spiracle, spiracle 2.3 from base of spinnerets. Palp: tibia as long as cymbium; tegulum oval; spermatic duct “Ushaped” ventrally; short distal tegular projection, close to the embolus apex; subdistal median apophysis, inverted “S-shaped”; embolus with median prolateral origin, thin, with broad base, slightly curved ( Figs 1C , 2A , 3B, C ).

Female (Paralectotype BMNH). Coloration as in male, except dorsal area of the abdomen lighter ( Fig. 1B ). Total length 9.2, carapace length 3.8, width 2.8. Clypeus height 0.16. Eye diameters and interdistances: AME 0.11, ALE 0.20, PME 0.16, PLE 0.18; AME–AME 0.07, AME–ALE 0.06, PME–PME 0.22, PME–PLE 0.18, ALE–PLE 0.06. Chelicerae 1.67 long, five promarginal teeth, eight retromarginal denticles. Leg measurements: leg I—femur 4.8/ patella 1.55/ tibia 5.1/ metatarsus 4.1/ tarsus 2.1/ total 17.65; II—4.3/ 1.4/ 4.5/ 3.85/ 1.7/ 15.75; III—3.1/ 1.2/ 2.4/ 3.1/ 1.0/ 10.8; IV—4.6/ 1.4/ 4.1/ 5.0/ 5.0/ 1.3/ 16.4. Leg spination: I—tibia v2-2-0, p1-0-0, r1-0-0, metatarsus v2-0-0, p0, r0; II—tibia v2-2-0, p1-1-1, r0, metatarsus v2-0-0, p1-0-0, r0; III—IV—tibia v2-2-2, p1-1-1-, r1-1-1, metatarsos v2-2-2, p1-1-1, r1-1-1. Abdomen: length 5.7, epigastric furrow 1.55 from tracheal spiracle, spiracle 2.25 from base of spinnerets. Epigynum: lateral borders narrow, not distally joined; internally with long, coiled copulatory ducts; seminal receptacles in the proximal area of the copulatory ducts; oval spermathecae; fertilization ducts as long as the spermathecae, curved and basally inserted ( Figs 1F, G , 2C, D , 3F ).

Variation. Males (n=10): total length 7.4–9.2; carapace length 2.8–3.8; femur I length 3.8–5.2. Females (n=10): total length: 8.5–9.7; carapace length 3.3–3.6; femur I length 4.2–4.8.

Distribution. Northeast to southern Brazil, West Paraguay and Northeastern Argentina ( Fig. 27C ).

Figures

FIGURE 1A–G. Patrera procera (Keyserling, 1981). Male (IBSP 138902): A habitus, dorsal view; C left palp, ventral view; D same, retrolateral view; E same, dorsal view. Female (IBSP 138902): B habitus, dorsal view; F epigynum, ventral view; G epigynum, dorsal view. Abbreviations: A, atrium; dTA, dorsal tibial apophysis; E, embolus; LB, lateral borders; MA, median apophysis; RTA, retrolateral tibial apophysis; T, tegulum. Scale bars: A, B 3.5mm; C 0.6mm; D 0.73mm; E 0.66mm; F 0.4mm; G 0.25mm.

FIGURE 2A–D. Patrera procera (Keyserling, 1891). Male (MCN 10965): A left palp, ventral view; B same, retrolateral view. Female (MCN 3172): C epigynum, ventral view (arrow indicating the excavation at the lateral borders); D same, dorsal view. Abbreviations:A, atrium; CD, copulatory ducts; dTA, dorsal tibial apophysis; E, embolus; FD, fertilization ducts; H, hood; LB, lateral borders; MA, median apophysis; RTA, retrolateral tibial apophysis; S, spermathecae; SR, seminal receptacles; T, tegulum. Scale bars: A–D 0.25mm.

FIGURE 3A–F. Patrera procera (Keyserling, 1891). Male (IBSP 139486):A chelicerae, ventral view; B left palp, ventral view; C same, distal view; D same, ventral view; E same, retrolateral view. Female (IBSP 138902): F epigynum, ventral view (arrow indicating the excavation at the lateral borders). Abbreviations: A, atrium; CO, copulatory opening; dTA, dorsal tibial apophysis; E, embolus; LB, lateral borders; MA, median tibial apophysis; RTA, retrolateral tibial apophysis; T, tegulum. Scale bars: A 0.3mm; B 0.4mm; C, F 0.2mm; D, E 0.1mm.

FIGURE 27A–F. Distribution map of Patrera species.