Clavatula ’ santhai Kovács & Vicián, 2021

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 29

‘ Clavatula ’ santhai Kovács & Vicián, 2021

Figs 11G 1 –G 2 , 3N , 5 , 6

Clavatula santhai n. sp. — Kovács & Vicián 2021: 144, pl. 3, figs 3–6.

Type material. Holotype: HNHM, PAL 2021.20 .1, SL: 37.0 mm, MD: 13.5 mm, Letkés ( Hungary), illustrated in Kovács & Vicián (2021, pl. 3, figs 5–6), figs 11G 1 –G 2, 3N.

Revised description. Shell medium-sized, moderately slender fusiform with high spire; apical angle ~38°. Protoconch not preserved. Teleoconch of ten whorls. Earliest spire whorls with broad, smooth subsutural cord and row of suprasutural beads, separated by narrow spiral groove.Abapically, mid-portion widens and becomes concave, with axial riblets ending at the swollen suprasutural row of beads. Later whorls subcylindrical, subsutural collar weakly developed bearing small, pointed tubercles just below suture on last two whorls, weak central concavity with delicate spiral threads in some specimens, suprasutural beads strengthen to form axially elongated opisthocline tubercles, partly covered by succeeding whorl. Suture narrowly but deeply impressed, weakly undulating. Last whorl ~60% of the total height. Subsutural collar moderate in width, poorly delimited, with pointed tubercles adapically. Subsutural ramp, wide, weakly concave, delimited by tuberculose shoulder cord, convex below, weakly constricted at base. Siphonal fasciole moderately swollen, well delimited, twisted. Sculpture below ramp of seven primary cords. Shoulder cord tubercular, rest beaded. Peribasal cord slightly stronger, with secondary cords intercalated. Aperture moderately wide, ovate. Outer lip not thickened, smooth within. Siphonal canal moderately long, narrow. Anal canal moderately wide and deep, asymmetrically U-shaped, with apex mid-ramp. Columella excavated in upper third, straight below, slightly twisted at fasciole, smooth. Columellar and parietal callus thickened, sharply delimited, forming broad rim and narrow pseudumbilical chink.

Discussion. This species is very similar to ‘ Clavatula’ camillae ( Hoernes & Auinger, 1891) and differs mainly in its more slender outline, less concave whorls, weaker spiral sculpture on the base, which is also less constricted, and lacks the double perifasciolar cords seen exclusively in ‘ C ’ camillae .

Paleoenvironment. The occurrence at Letkés ( Hungary) suggests a preference for inner neritic environments in the vicinity of corals ( Kovács & Vicián, 2013).

Distribution in Central Paratethys. Badenian (middle Miocene): Pannonian Basin: Letkés ( Hungary) ( Kovács & Vicián 2021).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Descriptive terms used in descriptions and measurements to shell morphology, whorl profiles and anal sinus. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height.AL: aperture length.AW: aperture width.

FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici.

FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae.

FIGURE 6. Apertures of Paratethyan Clavatulidae (C. = Clavatula, ‘C.’ = ‘Clavatula’, G. = Granulatocincta, M. = Megaclavaltula, O. = Olegia).

FIGURE 11 A1–A3. Clavatula sorini nov. sp., holotype, NHMW 1865/0001/0224, Lăpugiu de Sus (Romania). B1–B3. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania).C1–C3. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania).D1–D3. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0042b, Lăpugiu de Sus (Romania), E1–E2. ‘Clavatula’ atatuerki nov. sp., holotype,. NHMW 1847/0058/1216, Seyithasan (Turkey). F1–F2. ‘Clavatula’ atatuerki nov. sp., paratype, NHMW 1847/0058/1217, Seyithasan (Turkey). G1–G2. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).