Clavatula ’ vitalisi Strausz, 1955

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 38-39

‘ Clavatula ’ vitalisi Strausz, 1955

Figs 14C 1 –C 2 , 5 , 6

* Clavatula interrupta vitalisi nov. f. — Strausz 1955: 202, pl. 12, figs 25–26.

Clavatula interrupta vitalisi Strausz, 1955 — Strausz 1966: 404, pl. 15, figs 2–3.

Clavatula interrupta vitalisi Strausz — Nicorici 1972: 69.

Clavatula vitalisi ( Strausz, 1955) [sic]— Landau et al. 2013: 288.

Type material. Holotype: M.105, Várpalota ( Hungary), SL: 57.8 mm, MD: 25.7 mm, illustrated in Strausz (1955, pl. 12, figs 25–26), stored in the Geological and Geophysical Institute of Hungary (Budapest), figs 14C 1 –C 2.

Revised description. Shell moderately large, stout biconic; apical angle 38°. Protoconch and earliest teleoconch whorls not preserved. Early teleoconch whorls flat-sided; sculpture abraded. Later whorls with moderate width, poorly delimited subsutural collar bearing low tubercles; concave mid-portion bearing comma-shaped axial riblets. Suprasutural cord with axially elongated tubercles. Entire shell surface covered by numerous secondary spiral cords also overriding tubercles. Dense spirals and prominent axial growth lines give surface slightly reticulated aspect. Suture narrowly incised, running below beads. Last whorl ~60% of total height. Subsutural collar of moderate width, weakly tubercular. Subsutural ramp narrow, concave. Shoulder inflated, delimited by row of axially elongated tubercles, mid-whorl weakly convex, weakly angled at base. Base moderately constricted, siphonal fasciole swollen, twisted; deep pseudoumbilicus. Sculpture below ramp of slightly strengthened tubercular shoulder cord, slightly strengthened peribasal and perifasciolar cords, with further cords of roughly alternate strength intercalated below shoulder. All cords faintly beaded. Aperture moderately wide, pyriform. Outer lip not thickened, smooth within. Anal sinus moderately wide and deep, symmetrically U-shaped, with apex mid-ramp. Siphonal canal moderately short, moderately narrow, deflected to left, unnotched. Columella excavated in upper third, straight below, smooth. Columellar and parietal callus strongly thickened, sharply delimited.

Discussion. ‘ Clavatula ’ vitalisi was established by Strausz (1955) as subspecies of Clavatula interrupta ( Brocchi, 1814) , but differs from that species distinctly in its stout biconic shape, the very prominent spiral cords and the short siphonal canal. Similarly, ‘ Clavatula ’ sophiae ( Hoernes & Auinger, 1891) differs in its more slender shape, longer siphonal canal, wider anal canal and presence of lirae within the outer lip. The morphologically closest species in the Paratethys Sea is ‘ Clavatula ’ hirmetzli Kovács & Vicián, 2021 , which has an even stouter outline.

Paleoenvironment. The assemblages from Várpalota ( Hungary) suggest inner neritic sandy bottoms ( Harzhauser et al. 2002).

Distribution in Central Paratethys. Badenian (middle Miocene): Pannonian Basin: Várpalota ( Hungary) ( Strausz 1966); Șimleu Basin: Tusa ( Romania) (Nicorci 1972).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Descriptive terms used in descriptions and measurements to shell morphology, whorl profiles and anal sinus. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height.AL: aperture length.AW: aperture width.

FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae.

FIGURE 6. Apertures of Paratethyan Clavatulidae (C. = Clavatula, ‘C.’ = ‘Clavatula’, G. = Granulatocincta, M. = Megaclavaltula, O. = Olegia).

FIGURE 14 A1–A3. ‘Clavatula’ olgae (Hoernes &Auinger, 1891), paralectotype, NHMW 1862/0001/0552a, Buituri (Romania). B1–B3. ‘Clavatula’ olgae (Hoernes & Auinger, 1891), lectotype, NHMW 1862/0001/0552, Buituri (Romania). C1–C2. ‘Clavatula’ vitalisi Strausz, 1955, holotype, M.105, Várpalota (Hungary). D1–D2. ‘Clavatula’ ariejansseni nov. sp., holotype, NHMW 1847/0058/1220, Seyithasan (Turkey). E1–E2. ‘Clavatula’ ariejansseni nov. sp., paratype, NHMW 1847/0058/1221, Seyithasan (Turkey).