Granulatocincta contorta ( Švagrovský, 1958 ) Harzhauser & Landau & Janssen, 2022

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 50

Granulatocincta contorta ( Švagrovský, 1958) nov. comb.

Figs 17F 1 –F 2 , G 1 –G 2 , 3Z , 5

* Clavatula (Clavatula) contorta n. sp. — Švagrovský 1958: 15, pl. 4, figs 3–7.

Clavatula (Clavatula) contorta Švagrovský —Švagrovský 1959: 223, pl. 8, fig. 6.

Clavatula (Clavatula) contorta Švagrovský —Švagrovský 1960a: 102, pl. 13, figs 2–3.

Type material. Holotype: Z 3076, Kuzmice ( Slovakia), SL: 15.3 mm, MD: 6.9 mm, illustrated in Švagrovský (1958, pl. 4, fig. 5), stored in the Slovak National Museum (Bratislava, Slovakia), figs 17F 1 –F 2, 3Z. Paratypes: Z 3074, illustrated in Švagrovský (1958, pl. 4, figs, 3), figs 16G 1 –G 2; Z 3075, Z 3077, Z 3078, illustrated in Švagrovský (1958, pl. 4, figs, 4, 6, 7), stored in the Slovak National Museum (Bratislava, Slovakia).

Revised description. Shell small, solid, moderately broad buccinoid with conical spire; apical angle ~43°. Protoconch paucispiral of one whorl. Teleoconch of up to nine whorls. Early teleoconch whorls flat-sided, bearing weakly opisthocline axial ribs with tripartite sculpture; two beaded adsutural spiral cords, separated by narrower beaded central cord. Subsutural cord becoming coarsely tuberculose, central and suprasutural band with smaller tubercles, further finely granular secondary spirals intercalated. Suture narrowly impressed, undulating. Last whorl 65% of total height, subcylindrical to weakly ovoid; subsutural collar broad, poorly delimited, weakly tuberculose. Subsutural ramp weakly concave, narrow, with beaded cord running at mid-ramp. Shoulder poorly defined by beaded cord, weakly convex below, weakly angular at peribasal cord. Base weakly constricted. Siphonal fasciole moderately swollen, twisted. Peribasal and perifasciolar cords weakly tuberculose, the tubercles aligned axially to form indistinct axial riblets, with numerous secondary cords of irregular strength intercalated over entire whorl. Aperture wide, ovate. Outer lip not thickened, smooth within. Anal sinus wide, deep, symmetrically U-shaped. Apex coinciding with row of beads mid-ramp. Siphonal canal moderately short, shallowly notched at tip. Columella weakly excavated in upper third, straight below, smooth. Columellar and parietal callus strongly thickened, sharply delimited, forming broad callus rim.

Discussion. Granulatocincta contorta ( Švagrovský, 1958) is slightly reminiscent of Granulatocincta callim nov. sp. The general mode of sculpture is similar in the two species, but G. callim differs in its more subcylindrical last whorl and the sculpture of G. callim is distinctly more crowded and stronger.

The Tortonian, Proto-Mediterranean Granulatocincta sotterii ( Michelotti, 1847) is similar, but differs in its more crowded sculpture, much more prominent beads and more convex last whorl (see Montanaro 1937: 117, pl. 5, figs 2–18; Ruggieri & Davoli 1984, pl. 4, figs 1, 6, 13–14, 18–19, Davoli 1990, pl. 8, figs 2–8). Granulatocincta pelliscrocodili nov. sp., from the Serravallian of Turkey, might be a related species, but differs from G. contorta in its stout shape, the even weaker beads and the delicate spiral sculpture, which overruns the beads.

Paleoenvironment. Inner neritic environments ( Švagrovský 1960b).

Distribution in Central Paratethys. Late Badenian (middle Miocene): East Slovakian Basin: Kuzmice ( Slovakia) ( Švagrovský 1958, 1959, 1960a).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Descriptive terms used in descriptions and measurements to shell morphology, whorl profiles and anal sinus. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height.AL: aperture length.AW: aperture width.

FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici.

FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae.

FIGURE 17 A1–A3. Granulatocincta callim nov. sp., holotype, NHMW 1866/0001/0549, Wiesen (Austria). B1–B3. Granulatocincta callim nov.sp.,paralectotype,NHMW 1846/0037/0351b, Wiesen (Austria).C1–C3. Granulatocincta callim nov. sp., paralectotype, NHMW 1846/0037/0351, Wiesen (Austria). D1–D2. Granulatocincta callim nov. sp., paralectotype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). E. Granulatocincta carasiensis (Jekelius, 1944), MNG P– 4965, Soceni (Romania). F1–F2. Granulatocincta contorta (Švagrovský, 1958), holotype, Z 3076, Kuzmice (Slovakia). G1–G2. Granulatocincta contorta (Švagrovský, 1958), paratype, SNMB Z 3074, Kuzmice (Slovakia). H1–H3. Granulatocincta pelliscrocodili nov. sp., holotype, RGM 776 907, Gödet Creek near Tilkikaya (Turkey). I1–I2. Granulatocincta pelliscrocodili nov. sp., paratype, RGM 776 906, southwest bank of Gödet Creek near Tilkikaya (Turkey). J1–J3. Granulatocincta nataliae (Hoernes & Auinger, 1891), holotype, NHMW 1855/0035/0021, Belgrade/Rakovica (Serbia).