Granulatocincta theoderichi, Harzhauser & Landau & Janssen, 2022

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 58-60

Granulatocincta theoderichi nov. sp.

Figs 19E 1 –E 3 , F 1 –F 3 , G 1 –G 3 , H 1 –H 3, 3 Aa, 5, 6

Type material. Holotype: NHMW 2016 /0026/0320a, SL: 34.4 mm, MD: 13.0 mm, Ritzing ( Austria), figs 19E 1 –E 3 . Paratypes: NHMW 2016 /0026/0320b, SL: 33.1 mm , MD: 13.2 mm, Ritzing ( Austria) , figs 19F 1 –F 2, 3Aa; NHMW 1848 /0003/0022a, SL: 25.2 mm , MD: 10.3 mm, Ritzing ( Austria) , figs 19G 1 –G 3; NHMW 1848 /0003/0022b, SL: 20.3 mm , MD: 8.7 mm, Ritzing ( Austria) , figs 19H 1 –H 3; NHMW 2013 /0297/0012, 3 spec., Ritzing ( Austria) .

Type locality. Ritzing ( Austria); Oberpullendorf Basin .

Type stratum. Silty sand of the Ritzing Formation.

Age. Middle Miocene, late Badenian (Serravallian).

Etymology. Referring to Theodoric the Great (454–526), king of the Ostrogoth, who was born in the area, which today belongs to the Burgenland province in Austria.

Diagnosis. Granulatocincta species of medium size, solid, slender fusiform shell, strongly coronate, weakly gradate spire, tripartite early spire whorls. Last whorl with broad tuberculate subsutural collar. Narrow subsutural ramp bearing fine spiral cord. About 12 cords below ramp bearing coarse beaded sculpture.

Description. Shell medium-sized, solid, fusiform, with strongly coronate, weakly gradate spire; apical angle ~40–45°. Protoconch not preserved. Teleoconch of nine whorls. Earliest teleoconch whorls flat-sided with tripartite sculpture; indistinctly beaded subsutural cord, weakly beaded central cord. Coarsely beaded suprasutural cord. Sculpture changing gradually from fourth whorl; prominent, swollen subsutural collar with pointed tubercles (14– 16 on last whorl) developed on adapical half of collar, central row of beads coalesce to form smooth, narrow central cord. Suprasutural row of beads further strengthens, overrun by one or two weak, narrow cords. Whorl profile on later whorls subcylindrical with narrow central concavity filled by central cord. Suture narrowly incised, undulating below beads. Last whorl ~60% of total height. Subsutural collar broad, swollen, sharply delimited, bearing small, pointed tubercles. Subsutural ramp narrow, concave, bearing single smooth, narrow cord, shoulder delimited by beaded cord, broadly concave below, weakly constricted at base. Siphonal fasciole strongly swollen, twisted. Sculpture below ramp of about 12 beaded spiral cords, separated by narrow grooves at mid-whorl, becoming wider spaced towards fasciole. Narrow, unbeaded cords over fasciole. Prominent axial growth lines. Aperture moderately narrow, pyriform. Outer lip not thickened, smooth within. Anal sinus moderately narrow and deep, symmetrically Ushaped, with apex on central cord. Siphonal canal moderately long, moderately narrow, straight, shallowly notched at tip. Columella strongly excavated in upper third, straight below, smooth. Columellar and parietal callus strongly thickened, sharply delimited, forming broad callus rim and pseudumbilical chink.

Discussion. This species becomes gradually more elongate due to the ontogentic increase of the height of the last whorl. It is quite variable in sculpture, especially concerning the strength of the tubercles. This variability might be an expression of the unstable ecological conditions in the intertidal to shallow sublittoral zone.

Granulatocincta theoderichi is unique within the Paratethyan Clavatulidae due to its coarse sculpture. Olegia agathae ( Hoernes & Auinger, 1891) is superficially similar, but differs in its less protruding subsutural collar and much wider central concavity of the spire whorls. In addition, O. agathae has fewer spiral cords on the base. The same features distinguish Olegia mandici nov. sp. from G. theoderichi . Olegia szokolyensis ( Strausz, 1960) differs in its more conical spire whorls and the much weaker spiral sculpture on the base. A placement in Olegia is excluded based on the tripartite sculpture of early teleoconch whorls.

Paleoenvironment. Coastal, intertidal to inner neritic sandy bottoms in a restricted lagoon based on lithology and molluscan assemblages (own data).

Distribution in Central Paratethys. Badenian (middle Miocene): Oberpullendorf Basin: Ritzing ( Austria).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici.

FIGURE 19 A1–A3. Granulatocincta schreibersi (Hörnes, 1854), lectotype, NHMW 1846/0037/0315, Steinebrunn (Austria). B1–B3. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW 2021/0130/0001, Enzesfeld (Austria). C1–C3. Granulatocincta schreibersi (Hörnes, 1854), NHMW 1863/0015/1131, Gainfarn (Austria). D1–D3. Granulatocincta schreibersi (Hörnes, 1854), NHMW 2021/0126/0007, Gainfarn (Austria). E1–E3. Granulatocincta theoderichi nov. sp., holotype, NHMW 2016/0026/0320, Ritzing (Austria). F1–F3. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). G1–G3. Granulatocincta theoderichi nov. sp., paratype, NHMW 1848/0003/0022a, Ritzing (Austria). H1–H3. Granulatocincta theoderichi nov. sp., paratype, NHMW 1848/0003/0022b, Ritzing (Austria).