Olegia mandici, Harzhauser & Landau & Janssen, 2022

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 85-86

Olegia mandici nov. sp.

Figs 26A 1 –A 2 , B 1 –B 2 , C 1 –C 2 , D 1 –D 2 , 3 Ac, 5, 7

Pleurotoma doderleini Hörn. — Pilar 1874: 216, pl. 2, figs 20–21 [non Olegia doderleini ( Hörnes, 1854) ].

Clavatula doderleini brigittae Hoernes & Auinger, 1892 — Strausz 1966: 410, pl. 16, fig. 15, pl. 17, fig. 1 [non Olegia brigittae Hörnes & Auinger, 1891 ]

Type material. Holotype: NHMW 2021 /0140/0001, SL: 27.8 mm, MD: 11.2 mm, Bekića vrelo, Čremušnica at Donji Kirin ( Croatia), figs 26A 1 –A 2 . Paratypes: NHMW 2021 /0140/0002, SL: 25.3 mm , MD: 10.5 mm, Bekića vrelo, Čremušnica at Donji Kirin ( Croatia) , figs 26B 1 –B 2; NHMW 1872 /0031/0008a, SL: 23.4 mm , MD: 9.4 mm, Bekića vrelo, Čremušnica at Donji Kirin ( Croatia) , figs 26C 1 –C 3; NHMW 1872 /0031/0008b, SL: 19.4 mm , MD: 8.4 mm, Bekića vrelo, Čremušnica at Donji Kirin ( Croatia) , figs 26D 1 –D 3; NHMW 1872 /0031/0008c, SL: 27.9 mm , MD: 13.5 mm, Bekića vrelo, Čremušnica at Donji Kirin ( Croatia) , fig. 3Ac; NHMW 1872 /0031/0008, 4 spec., Bekića vrelo, Čremušnica at Donji Kirin ( Croatia) .

Additional studied material. NHMW 1858 /0015/0148, 1 spec., Totterfeld at Hartberg ( Austria); GBA 1875/006/0011, Hafnertal at Lichtenwald ( Austria), illustrated in Hoernes (1875b, pl. 2, fig. 4) .

Type locality. Bekića vrelo, Čremušnica at Donji Kirin ( Croatia), Glina Basin .

Type stratum. Badenian siliciclastic deposits (no formation name available).

Age. Middle Miocene, Sarmatian (late Serravallian).

Etymology. In honor of Oleg Mandic (paleontologist at the NHMW and friend of the first author) in recognition of his contributions on Paratethyan stratigraphy.

Diagnosis. Olegia species of medium size, solid moderately slender fusiform shell, with strong sculpture of pointed tubercles on subsutural collar and rounded tubercles on suprasutural cord separated by wide, smooth concave mid-whorl portion. Four tubercular spiral cords on last whorl with secondary and tertiary spiral threads intercalated.

Description. Shell small, solid, moderately slender fusiform with conical spire; apical angle ~35°. Protoconch not preserved. Teleoconch of at least eight whorls. Early teleoconch whorls flat-sided with bipartite sculpture; welldefined, smooth subsutural cord, smooth mid-whorl portion. Suprasutural cord bearing large, rounded, slightly opisthocline tubercles. Later whorls with moderate width subsutural collar bearing pointed tubercles, concave midwhorl with comma-shaped riblets, and suprasutural row of larger orthocline tubercles. Entire surface covered by secondary and tertiary cords. Suture narrowly but deeply incised, weakly undulating, running just below abapical row of tubercles. Last whorl ~ 65% of total height; subsutural collar well-defined bearing about 14 small, sharp tubercles, subsutural ramp concave, delimited by tubercular shoulder cord, convex below, weakly to moderately constricted at base. Siphonal fasciole swollen, strongly twisted. Sculpture of four tubercular primary spiral cords. Adapical cord delimiting shoulder strongest, second cord at mid-whorl delimiting periphery, third peribasal, fourth perifasciolar, one further slightly weaker primary cord intercalated between shoulder and peripheral primaries and one or two on base between peribasal and perifasciolar cords. Threads of tertiary strength cover entire surface. Aperture moderately wide, ovate. Outer lip not thickened, smooth within. Anal sinus deep, moderately wide, symmetrically U-shaped, with apex just below collar. Siphonal canal moderately long and narrow, deflected to the left, shallowly notched at tip. Columella weakly excavated, twisted at fasciole, smooth. Columellar and parietal callus strongly thickened, sharply delimited, forming very broad callus rim and narrow pseudumbilical chink.

Discussion. Olegia mandici nov. sp. is reminiscent of Olegia rumana ( Simionescu & Barbu, 1940) and might represent an endemic offshoot of that species. It differs from O. rumana in its larger size, the less numerous but larger beads at the abapical suture of early teleoconch whorls, the regular sculpture and more pointed tubercles, the wider spiral cords on the last whorl bearing larger tubercles, and the presence of secondary and tertiary spiral sculpture. The early Miocene Olegia agathae ( Hoernes & Auinger, 1891) might be another closely related species, which differs especially in its smaller size and the higher, weakly constricted base. Olegia szokolyensis ( Strausz, 1960) differs in its lower spire, the subcylindrical last whorl and the much weaker spiral sculpture on the base. This specimen is either a misplaced Sarmatian specimen or the only known Badenian representative.

Distribution in Central Paratethys. Sarmatian (middle Miocene): Styrian Basin: Totterfeld at Hartberg ( Austria) (hoc opus), Hafnertal at Lichtenwald ( Austria) ( Hoernes 1875b); Glina Basin: Bekića vrelo, Čremušnica at Donji Kirin ( Croatia) (hoc opus). Strausz (1966) described a specimen from the Badenian of Budapest as Clavatula doderleini brigittae , which seems to represent O. mandici based on its spiny sculpture and spiral sculpture.

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Descriptive terms used in descriptions and measurements to shell morphology, whorl profiles and anal sinus. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height.AL: aperture length.AW: aperture width.

FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici.

FIGURE 26 A1–A2. Olegia mandici nov. sp., holotype, NHMW 2021/0140/0001, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). B1–B2. Olegia mandici nov. sp., paratype, NHMW 2021/0140/0002, Bekića vrelo, Čremušnica (Croatia). C1–C2. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). D1–D2. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008b, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). E1–E2. Olegia rumana (Simionescu & Barbu, 1940), ZNG PAN A-I-50/467, Chlebów (Poland). F1–F2. Olegia rumana (Simionescu & Barbu, 1940), ZNG PAN A-I-50/468, Chlebów (Poland). G1–G2. Olegia rumana (Simionescu & Barbu, 1940), ZNG PAN A-I-50/468, Chlebów (Poland).