Hurites, Tian & Huang & Jia, 2023

Tian, Mingyi, Huang, Sunbin & Jia, Xinyang, 2023, A contribution to cavernicolous beetle diversity of South China Karst: eight new genera and fourteen new species (Coleoptera: Carabidae: Trechini), Zootaxa 5243 (1), pp. 1-66 : 28

Hurites n. gen.

Type species: Hurites yunxianicus n. sp. (Yunxian Dong, Shaoyang , Hunan)

Generic characteristics. Large-sized for Cimmeritodes -like species, whole pubescent, anophthalmic, fore part (head plus pronotum) evidently shorter than elytra; head elongate, nearly parallel-sided, right mandible tridentate with teeth blunt; frontal furrows not parallel-sided, two pairs of supraorbital setiferous pores present, mentum and submentum fused; antennae rather short, extending to apical 1/4 of elytra; pronotum subquadrate, fore lateromarginal angles protruding, hind angles nearly rectangular, presence of two pairs of latero-marginal setae; elytra elongate, sides subserrate and ciliate throughout; surface densely punctate, striae faint and more or less traceable, intervals slightly convex; presence of two dorsal pores and the preapical pores on each elytron; the humeral group of marginal umbilicate pores aggregated, the 1 st and 2 nd pores close to marginal gutter; the 5 th and 6 th pores of the middle group very widely spaced, making 5 th much closer to 4 th than to 6 th; the 1 st and 2 nd protarsomeres widened, and denticulate inwardly at apices in male; male genitalia weakly-sclerotized, thin and elongate, presence of a moderately sized sagittal aileron, parameres much shorter than median lobe.

Remarks. Hurites n. gen. is probably a member of Cimmeritodes Deuve, 1996 complex, in particular somewhat similar to the subgenus Shimenrites Deuve & Tian, 2017, which is also occurring in Hunan Province ( Deuve & Tian 2017). However, it differs from the latter in: (1) mentum and submentum fused in Hurites n. gen., instead of separated in Shimenrites; (2) median group (5 th and 6 th pores) of the marginal umbilicate series extraordinarily removed each other in Hurites n. gen., instead of close to each other in Shimenrites; (3) the 1 st and 2 nd protarsomeres of male modified in Hurites n. gen., but both are simple in male in Shimenrites.

Regarding the fact that many subgenera of Cimmeritodes should be treated as independent genera according to phylogenetic analysis based on molecular data, we would like to propose Hurites as a new genus rather than a new subgenus.

Etymology. “Hu+ rites”, “Hu” means Hunan Province, indicating the distribution area of this new genus. Gender masculine.

Range. China (Hunan) ( Figure 1l ).

Figures

FIGURE 1. Map indicating the locality caves of the new genera and species. a. Xianren Dong; b. Shisanya Dong; c. Da Dong, Tuoluo; d. Da Dong, Bali; e. Weixiao Tiankeng; f. Laba Dong; g. Gui Dong; h. Suolue Cave I; i. Xiannü Dong; j. Jiudu Cave I; k. Xiannü Dong; l. Yunxian Dong; m. Zhongguo Dong; n. Feixian Dong.