Haemaphysalis rugosa Santos Dias, 1956b

Guglielmone, Alberto A., Nava, Santiago & Robbins, Richard G., 2023, Geographic distribution of the hard ticks (Acari: Ixodida: Ixodidae) of the world by countries and territories, Zootaxa 5251 (1), pp. 1-274 : 98

140. Haemaphysalis rugosa Santos Dias, 1956b .

Afrotropical: 1) Benin, 2) Burkina Faso, 3) Central African Republic, 4) Democratic Republic of the Congo, 5) Ghana, 6) Ivory Coast, 7) Senegal, 8) South Sudan, 9) Uganda ( Hoogstraal & El Kammah 1972, Matthysse & Colbo 1987, Kolonin 2009, Uilenberg et al. 2013, Sylla et al. 2018).

Haemaphysalis rugosa was originally identified as Haemaphysalis aciculifer by Nuttall & Warburton (1915). Later, Santos Dias (1956b) gave subspecific status to a specimen of Haemaphysalis aciculifer that he called Haemaphysalis aciculifer rugosa , which was, in turn, given specific status by Hoogstraal & El Kammah (1972). The latter authors emphasized that Haemaphysalis rugosa had been extensively confused with Haemaphysalis aciculifer , and the range of Haemaphysalis rugosa is based on Hoogstraal & El Kammah (1972) and subsequent publications.

Ntiamoa-Baidu et al. (2004) recognized the presence of Haemaphysalis aciculifer but not Haemaphysalis rugosa in Ghana, but this would constitute an error because the holotype of Haemaphysalis rugosa was collected in Ghana ( Keirans 1985b). ElGhali & Hassan (2012) did not recognize the presence of Haemaphysalis rugosa in South Sudan, but that country is here included within the range of this tick.