Dima florinensis Platia, 2012

Mertlik, Josef, Németh, Tamás & Kundrata, Robin, 2017, Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula, Zootaxa 4220 (1), pp. 1-63 : 16-17

Dima florinensis Platia, 2012

Figs 31–34 , 129 , 185 , 244 .

Dima florinensis Platia, 2012: 201 .

Type depository. Holotype, ♀ (PCGP).

Type locality. Greece: Flórina, Varnous Mts., A. Triados (40°47′32.1''N, 21°13′35.4''E), 1285 m.

New material. GREECE: distr. Flórina, Varnoús Mts., Pisoderi, Agia Triada env., 1410 m (40°47'32.1"N, 21°13'35.4"E), 27. V.2013, 2 ♂♂, J. Mertlik et V. Dušánek leg ( PCJM, PCVD) ; distr. Flórina, Varnoús Mts., Vigla pass, beechwood, 1565 m (40°46'20.80"N, 21°15'55.25"E), 27. V.2013, 4 ♂♂, 4 ♀♀, J. Mertlik leg. (PCJM); dtto, 3 ♂♂, 6 ♀♀, V. Dušánek leg. (PCVD); dtto, 4 ♂♂, 2 ♀♀, P. Brůha leg. ( PCPB); Makedonia, pref. Florina, Mt. Vernon , 2 km W Alona , beech forest, 40°46'52.97"N, 21°16'33.12"E, 1400 m, singled at night, 24 . VI.2012, 2 ♂♂, 5 ♀♀, leg. A. Kotán, A. Márkus, P. Nemes & T. Németh (PCRK); dtto, 1 ♂, 1 ♀ (HNHM); dtto, 18. V.2013, 2 ♀♀, leg. A. Kotán, P. Nemes, Gy. Németh, T. Németh & V. Szénási (HNHM); dtto, 2 ♂♂, 5 ♀♀ ( PCRK); prov. Makedonia, distr. Florina, Mt. Vernon , 2 km W Pisoderi ski center, 40°47'20.16"N, 21°13'29.51"E, 1330 m, beech forest, singled at night, 18 . V.2013, 2 ♂♂, 2 ♀♀, leg. Kotán, P. Nemes, Gy. Németh, T. Németh & V. Szénási (HNHM); dtto, 3 ♂♂, 3 ♀♀ (PCRK); MACEDONIA: Galičica, northern part [sev. str.], IX. 1926, 1 ♀, Dr. Rambousek leg. ( NMPC) ; Baba planina, Niže Pole – Golemo jezero, 1500–2000 m, 18. VII.2006, 1 ♀, V. Zieris leg. (PCJM); dtto, 14.–15. VII.2010, 1 ♀, V. Zieris leg. ( PCTS); Baba Mts., Malovište, Šemnica valley , 1600–1800 m, 11.–12 . VI.2008, 1 ♂, L. Blažej leg. ( PCPB); distr. Bitola, Baba plateau, 1.7 km S of Nižepole, beechwood, 1400 m (40°58'47.54"N, 21°14'42.92"E), 26 . V.2014, 3 ♂♂, 1 ♀, J. Mertlik leg. (PCJM); dtto, 2 ♂♂, 3 ♀♀, B. Zbuzek leg. (PCBZ); dtto, 3 ♂♂, 1 ♀, P. Brůha leg. ( PCPB); distr. Bitola, Baba plateau, Nižepole – ski centre env., beechwood, 1375 m (40°58'52.22"N, 21°15'13.82"E), 26 . V.2014, 1 ♂, 2 ♀♀, J. Mertlik leg. (PCJM); dtto, 8 ♂♂, 2 ♀♀, B. Zbuzek leg. (PCBZ); dtto, 3 ♂♂, P. Brůha leg. ( PCPB); Nizhepole reg., NP Pelister, 2.5 km S Nizhepole, mixed forest, swept and singled at night, 40°59'3.65"N, 21°14'37.24"E, 6 . VI.2014, 2 ♂♂, 4 ♀♀, leg. V. Dušánek, R. Kundrata & T. Németh (HNHM); dtto, 27 ♂♂, 11 ♀♀ (PCRK); dtto, 18 ♂♂, 15 ♀♀, V. Dušánek leg. ( PCVD); Nizhepole reg., NP Pelister, 3 km S Nizhepole, 40°58'52.5"N, 21°15'16.2"E, 6 . VI.2014, 1 ♂, leg. V. Dušánek, R. Kundrata & T. Németh ( PCRK); Pelagonia region, Pelister Mts., Nižepole , forest brook below the ski station, N40°58.889', E21°15.246', 1370 m, 25 . VI.2014, 1 ♂, leg. Tibor Kovács (MMGY).

Description of male. Like female, but with relatively longer antennae, surpassing the hind angles of pronotum by about three antennomeres (about one in females), and pronotum slightly narrower than in females ( Fig. 32 ). Aedeagus as in Fig. 185 .

Diagnosis. This species belongs to the group of species with a more convex, suboval-shaped body and shiny elytral intervals with very sparsely distributed thin setae ( Figs 31–34 ). Dima florinensis is a medium-sized species (body length: 11.0–13.0 mm) with long, almost tangled, semi-erect pubescence on the head and pronotal sides (typical for this species; Fig. 129 ), short antennae and by the distinct double punctation of pronotum ( Fig. 129 ). It is very like D. macedonica by the habitus, double punctation of pronotum, and shape and length of antennae. These species differ in the scutellum ( D. florinensis has a scutellum on the same plane as the elytra in lateral view; D. macedonica has a scutellum raised above the plane of the elytra; Fig. 164 ) and pubescence of the pronotal sides ( D. macedonica has pubescence in anterior third of the pronotal sides about 0.5 times as long as in the posterior half; Figs 141–143 ). D. raineri differs from D. florinensis by the decumbent pubescence in the first half of pronotal sides and double punctation of pronotum being less distinct, with punctures more similar in size to each other ( Figs 154– 155 ), and D. vailatii by the denser pubescence of the elytra and the shorter pubescence of pronotal sides ( Figs 109– 113 , 158–160 ). The sclerotized spines of bursa copulatrix as in Fig. 228 .

Intraspecific variability. There is an intraspecific variability in the length of the apical lobe of paramera in D. florinensis (similarly as in D. macedonica ). The holotype is darker than remaining specimens, which are reddishbrown ( Figs 31–34 ). There is a variability in the punctation density on the pronotum; most specimens have densely punctate pronotum, but some specimens from Nizhepole region have sparser punctures. The female collected in the northern slope of Galičica is paler and has much sparser pronotal punctation than the remaining specimens of this species ( Fig. 34 ).

Distribution. Greece (Varnous Mts., Mt. Verno), Macedonia (Baba Mts., Galičica Mts.) ( Fig. 244 ). First record for Macedonia.

Figures

FIGURE 244. The distribution of Dima species in Albania, Bulgaria, Macedonia and in the northern Greece.

FIGURES 30 – 44. Habitus images of Dima species. 30 — D. fialai sp. nov., paratype, female; D. florinensis Platia, 2012, 31 — holotype, female, 32 — non-type male from Varnous Mts., 33 — non-type female from Varnous Mts., 34 — non-type female from Galičica Mts.; D. fthiotidensis Schimmel & Platia, 2008, 35 — paratype, female, 36 — holotype of D. fokidensis Schimmel & Platia, 2008, female, 37 — holotype of D. scutellaris Platia, 2010, female, 38 — non-type male from Kallidromo Mts., 39 — nontype male from Vardoussia Mts., 40 — non-type female from Vardoussia Mts., 41 — non-type female from Vardoussia Mts.; D. giachinoi Schimmel & Platia 2008, 42 — paratype, female, 43 — non-type specimen, male, 44 — non-type specimen, female. Not to scale.

FIGURES 119 – 130. Pronotal details of Dima species. 119 — D. arndti Platia, 2013, male; 120 — D. assingi Schimmel & Platia, 2008, female; 121 — D. bruhai sp. nov., holotype, male; 122 — D. dalmatina Küster, 1844, male; 123 — D. dusaneki sp. nov., holotype, male; 124 — D. elateroides Charpentier, 1825, male; 125 — D. etoliensis Platia, 2012, holotype, female; 126 — D. evritaniensis Schimmel & Platia, 2008, paratype, female; 127 — D. evritaniensis, male; 128 — D. fialai sp. nov., holotype, male; 129 — D. florinensis Platia, 2012, female; 130 — D. fthiotidensis Schimmel & Platia, 2008, male. Not to scale.

FIGURES 182 – 191. Aedeagus images of Dima species. 182 — D. elateroides Charpentier, 1825, non-type specimen from Serbia; 183 — D. evritaniensis Schimmel & Platia, 2008, non-type specimen from Timfristos Mts.; 184 — D. fialai sp. nov., holotype; 185 — D. florinensis Platia, 2012, non-type specimen; 186 — D. fthiotidensis Schimmel & Platia, 2008, non-type specimen from Kallidromo Mts.; 187 — D. fthiotidensis, non-type specimen from Vardoussia Mts.; 188 — D. giachinoi Schimmel & Platia 2008, non-type specimen; 189 — Dima sp. from Mani; 190 — D. hladilorum Schimmel, 1987, non-type specimen; 191 — D. kozufensis, sp. nov., holotype. Not to scale.

FIGURES 163 – 171. Scutellum-shape of Dima species. 163 — D. bruhai sp. nov.; 164 — D. macedonica Schimmel, 1993; 165 — D. raineri Wurst, 1997. Elytral details of Dima species. 166 — D. fthiotidensis Schimmel & Platia, 2008; 167 — D. hirtipennis Platia, 2011; 168 — D. macedonica Schimmel, 1993; 169 — D. olympica Meischnigg, 1934; 170 — D. parnonensis Schimmel & Platia, 2008; 171 — D. pecoudi Fleutiaux, 1943. Not to scale.

FIGURES 131 – 142. Pronotal details of Dima species. 131 — D. fthiotidensis Schimmel & Platia, 2008, paratype, female; 132 — D. scutellaris Platia, 2010, holotype, female; 133 — D. fthiotidensis, male; 134 — D. fthiotidensis, holotype of D. fokidensis Schimmel & Platia, 2008; 135 — D. giachinoi Schimmel & Platia 2008, female; 136 — D. hirtipennis Platia, 2011, holotype, female; 137 — D. hladilorum Schimmel, 1987, male; 138 — D. kozufensis sp. nov., paratype, female; 139 — Dima lebenbaueri Schimmel & Platia 2008, holotype, male; 140 — Dima lebenbaueri, non-type specimen, female; 141 — D. macedonica Schimmel, 1993, holotype, female, 142 — D. macedonica, female paratype of D. hladilorum from Galičica Mts. Not to scale.

FIGURES 143 – 154. Pronotal details of Dima species. 143 — D. macedonica Schimmel, 1993, male; 144 — D. marvani Mertlik & Dusanek, 2006, holotype, male; 145 — D. moraveci Schimmel & Platia, 2008, holotype, female; 146 — D. neumanni Platia, 2013, female; 147 — D. olympica Meschnigg, 1934, male; 148 — D. orientalis sp. nov., holotype, male; 149 — D. parnonensis Schimmel & Platia, 2008, female; 150 — D. pecoudi Fleutiaux, 1944, male; 151 — D. pelikani sp. nov., holotype, male; 152 — D. pelionensis sp. nov., paratype, female, 153 — D. pelionensis sp. nov., paratype, female paratype of D. fokidensis; 154 — D. raineri Wurst, 1997, female. Not to scale.

FIGURES 155 – 162. Pronotal details of Dima species. 155 — D. raineri Wurst, 1997, holotype, male; 156 — D. schimmeli Platia & Gudenzi, 2009, female; 157 — D. schnitteri Platia, 2013, female; 158 — D. vailatii Schimmel & Platia 2008, holotype, male; 159 — D. vailatii, male; 160 — D. riesei Platia, 2010, holotype, male; 161 — D. vonickai sp. nov., holotype, male; 162 — D. zbuzeki sp. nov., holotype, male. Not to scale.

FIGURES 105 – 118. Habitus images of Dima species. Dima schnitteri Platia, 2013, 105 — holotype, male, 106 — paratype, female, 107 — non-type specimen, male, 108 — non-type specimen, female; D. vailatii Schimmel & Platia 2008, 109 — holotype, male, 110 — paratype, female, 111 — non-type specimen, male, 112 — non-type specimen, female, 113 — holotype of D. riesei Platia, 2010, male; D. vonickai sp. nov., 114 — holotype, male, 115 — paratype, female; D. zbuzeki sp. nov., 116 — holotype, male, 117 — paratype, female, 118 — paratype, male. Not to scale.

FIGURES 222 – 233. The sclerotized spines of bursa copulatrix of Dima species. 222 — D. arndti Platia, 2013, paratype; 223 — D. bruhai sp. nov., paratype; 224 — D. dusaneki sp. nov., paratype; 225 — D. elateroides, Charpentier, 1825, non-type specimen; 226 — D. etoliensis Platia, 2012, holotype; 227 — D. fialai sp. nov., paratype; 228 — D. florinensis Platia, 2012, holotype; 229 — D. fthiotidensis Schimmel & Platia, 2008, holotype of D. scutellaris Platia, 2010; 230 — D. giachinoi Schimmel & Platia 2008, paratype; 231 — D. hirtipennis Platia, 2011, holotype; 232 — D. kozufensis sp. nov., paratype; 233 — D. neumanni Platia, 2013, non-type specimen. Not to scale.