Dima dalmatina Küster, 1844

Mertlik, Josef, Németh, Tamás & Kundrata, Robin, 2017, Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula, Zootaxa 4220 (1), pp. 1-63 : 9-10

Dima dalmatina Küster, 1844

Figs 14–19 , 122 , 179–180 , 243 .

Dima dalmatina Küster, 1844: 13 ; Schwarz 1906: 247; Petri 1912: 196; Schenkling 1927: 419; Depoli 1928: 223; Fleutiaux 1943: 40; Novak 1952: 135; Cobos 1961: 109; Novak 1970: 21; Dajoz 1973: 105; Schimmel 1987: 42; Platia 1994: 121; Schimmel 1996: 138; Mertlik & Dusanek 2006: 151; Cate 2007: 183; Schimmel & Platia 2008: 566.

Type depository. Syntype, male (BMNH). Other type material unknown.

Type locality. Croatia: Omiš [Dalmatien: Almissa].

New material. BOSNIA AND HERZEGOVINA: Prenj planina, no date, 1 ex. [without head] ( PCGP) ; CROATIA: Zaostorog, Rilič [vápencové škrapy Riliče], 20. III.2000, 1 ♂, on the ground between stones [skalní lesostep, na zemi mezi kameny], R. Gabzdil leg. ( PCRG) ; Zaostorog , 22.– 24. III.2001, 1 ♀, P. Arpáš leg. ( PCJM); MONTENEGRO : Jugoslavia, Petrovac, 16. VI.1974, 1 ♀, J. Hladil leg. ( PCRS) ; Bar distr., Mt .. Vrsouta , Sutorman , mixed forest (42°9'18.32"N, 19°5'52.81"E), 1.–2. VI.2014, 2 ♀♀, singled at night, swept and beaten, V. Dušánek leg. ( PCVD) ; dtto, 1 ♂, 2 ♀♀, T. Németh leg. (PCRK), dtto, 2 ♀♀, Kundrata lgt. (PCRK); Dalmatien, Budva, coll. Reitter, 1 ♀ (HNHM); Dalmatia, coll. E. Frivaldszky, 1 ♂ (HNHM); Dalmatia, coll. Lichtn., 1 ♀ (HNHM).

Diagnosis. Dima dalmatina is a medium-sized species (body length: 11.5–13.5 mm) easily recognizable by its elongate body, relatively long antennae (surpassing hind angles by about 3.5 segments in males), shiny pronotum with sides with the short semi-erect setae along the whole length, elytra with short dense decumbent pubescence, and elytral interstices matt, microreticulated ( Figs 14–19 , 122 ). This species is very like D. marvani Mertlik & Dusanek, 2006 ( Figs 62–65 ). The typical representatives of D. dalmatina differ from the typical D. marvani by more distinctly defined keel between frons and clypeus, shorter antennae in males (about 55% of body length in D. dalmatina , more than 60 % in D. marvani ), wider pronotum (1.4 times wider than long in D. dalmatina , 1.3 times wider than long in D. marvani ), pronotal pubescence denser, near sides directed towards posterior angles ( Figs 122 , 144 ), and apical lobe of paramera shorter ( Figs 180 , 198 ).

Intraspecific variability. There are two distinct Dima populations in Rumija Mts. in Montenegro; one resembles more D. dalmatina and the second is more like D. marvani . However, they both differ from the typical representatives of respective species ( Figs 18–19 , 64–65 ). Both of these local forms have a darker body and slightly more convex and longer pronotum than most of the typical specimens. The specimens similar in most aspects to D. dalmatina have longer apical lobes of paramera like those found in D. marvani ( Fig. 179 ). Unfortunatelly, without larger series of both species collected from more localities we are unable to make any taxonomical conclusions and we keep the populations from Rumija Mts. in D. dalmatina and D. marvani .

Distribution. Bosnia and Herzegovina, Croatia, Montenegro ( Fig. 243 ). First record for Bosnia and Herzegovina.

Remarks. The syntype of D. dalmatina deposited in the BMNH, London, is clearly a male, but without genitalia neither inside the abdomen nor glued on a label ( Figs 14–15 ); there is a label "Abdomen seems to be empty, R. Schimmel 19 85".

A specimen from Greece (Florina, 1100 m, 11.VII.1981, Osella leg.), which is mentioned under D. dalmatina by Guglielmi & Platia (1985: 194), is probably an erroneously identified specimen. That specimen was identified by Dr. Guglielmi and should be deposited in the Verona museum (G. Platia, pers. comm.), however it is probably misplaced in the collection (R. Salmaso, pers. comm.). It might be conspecific with D. florinensis Platia, 2012 , which occurs in that area ( Platia 2012).

One specimen from Rumija Mts. recorded under D. dalmatina in Schimmel & Platia (2008) was redetermined as D. marvani .

Figures

FIGURE 243. The distribution of Dima species in the northern Balkan Peninsula and surrounding areas.

FIGURES 1 – 14. Habitus images of Dima species. Dima arndti Platia, 2013, 1 — holotype, male, 2 — paratype, female, 3 — nontype specimen, male, 4 — non-type specimen, female; 5 — Dima sp. from Crete, male; D. assingi Schimmel & Platia, 2008, 6 — holotype, male, 7 — holotype of D. peloponnensis Schimmel & Platia, 2008, male, 8 — non-type male from Kilini Mts., 9 — nontype female from Kilini Mts.; D. bruhai sp. nov., 10 — holotype, male, 11 — paratype, female, 12 — paratype, male, 13 — paratype, male; D. dalmatina Küster, 1844, 14 — syntype, male, lateral view. Not to scale.

FIGURES 15 – 29. Habitus images of Dima species. Dima dalmatina Küster, 1844, 15 — syntype, male, 16 — non-type male from Vosac, Croatia, 17 — non-type female from Vosac, Croatia, 18 — non-type male from Rumija Mts., Montenegro, 19 — nontype female from Rumija Mts., Montenegro; D. dusaneki sp. nov., 20 — holotype, male, 21 — paratype, female; D. elateroides Charpentier, 1825, 22 — non-type male from Serbia; 23 — non-type male from Slovenia, 24 — non-type female from Croatia; 25 — D. etoliensis Platia, 2012, holotype, female; D. evritaniensis Schimmel & Platia, 2008, 26 — paratype, female, Panetoliko Mts., 27 — non-type male from Timfristos Mts., 28 — non-type female from Timfristos Mts.; 29 — D. fialai sp. nov., holotype, male. Not to scale.

FIGURES 119 – 130. Pronotal details of Dima species. 119 — D. arndti Platia, 2013, male; 120 — D. assingi Schimmel & Platia, 2008, female; 121 — D. bruhai sp. nov., holotype, male; 122 — D. dalmatina Küster, 1844, male; 123 — D. dusaneki sp. nov., holotype, male; 124 — D. elateroides Charpentier, 1825, male; 125 — D. etoliensis Platia, 2012, holotype, female; 126 — D. evritaniensis Schimmel & Platia, 2008, paratype, female; 127 — D. evritaniensis, male; 128 — D. fialai sp. nov., holotype, male; 129 — D. florinensis Platia, 2012, female; 130 — D. fthiotidensis Schimmel & Platia, 2008, male. Not to scale.

FIGURES 172 – 181. Aedeagus images of Dima species. 172 — D. arndti Platia, 2013, holotype; 173 — D. arndti, non-type specimen; 174 — Dima sp. from Crete; 175 — D. assingi Schimmel & Platia, 2008, holotype; 176 — D. assingi, non-type specimen from Kilini Mts.; 177 — D. peloponnensis Schimmel & Platia 2008, holotype; 178 — D. bruhai sp. nov., holotype; 179 — D. dalmatina Küster, 1844, non-type specimen from Sutorman, Montenegro; 180 — D. dalmatina, non-type specimen from Vosec, Croatia; 181 — D. dusaneki sp. nov., holotype. Not to scale.

FIGURES 60 – 74. Habitus images of Dima species. Dima macedonica Schimmel, 1993, 60 — non-type male from Ehloec, 61 — female paratype of D. hladilorum Schimmel, 1987 from Galičica Mts.; D. marvani Mertlik & Dusanek, 2006, 62 — holotype, male, 63 — female paratype from Lovčen Mts., 64 — female paratype from Rumija Mts., 65 — non-type male from Rumija Mts.; 66 — D. moraveci Schimmel & Platia 2008, holotype, female; D. neumanni Platia, 2013, 67 — holotype, male, 68 — non-type male from Erimanthos Mts., 69 — non-type female from Erimanthos Mts., 70 — non-type male from Aroania Mts., 71 — non-type female from Aroania Mts., 72 — non-type female from Aroania Mts., 73 — non-type female from Panachaiko Mts., 74 — non-type female from Panachaiko Mts. Not to scale.

FIGURES 143 – 154. Pronotal details of Dima species. 143 — D. macedonica Schimmel, 1993, male; 144 — D. marvani Mertlik & Dusanek, 2006, holotype, male; 145 — D. moraveci Schimmel & Platia, 2008, holotype, female; 146 — D. neumanni Platia, 2013, female; 147 — D. olympica Meschnigg, 1934, male; 148 — D. orientalis sp. nov., holotype, male; 149 — D. parnonensis Schimmel & Platia, 2008, female; 150 — D. pecoudi Fleutiaux, 1944, male; 151 — D. pelikani sp. nov., holotype, male; 152 — D. pelionensis sp. nov., paratype, female, 153 — D. pelionensis sp. nov., paratype, female paratype of D. fokidensis; 154 — D. raineri Wurst, 1997, female. Not to scale.

FIGURES 192 – 201. Aedeagus images of Dima species. 192 — D. lebenbaueri Schimmel & Platia 2008, holotype; 193 — D. lebenbaueri, non-type specimen; 194 — D. macedonica Schimmel, 1993, holotype; 195 — D. macedonica, paratype; 196 — D. macedonica, non-type specimen from Galičica Mts.; 197 — D. macedonica, non-type specimen from Ehloec; 198 — D. marvani Mertlik & Dusanek, 2006, holotype; 199 — D. neumanni Platia, 2013, holotype; 200 — D. neumanni, non-type specimen from Erimanthos Mts.; 201 — D. neumanni, non-type specimen from Aroania Mts. Not to scale.