Dima dusaneki, Mertlik, Josef, Németh, Tamás & Kundrata, Robin, 2017

Mertlik, Josef, Németh, Tamás & Kundrata, Robin, 2017, Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula, Zootaxa 4220 (1), pp. 1-63 : 10-11

Dima dusaneki sp. nov.

Figs 20–21 , 123 , 181 , 224 , 244 .

Type material. Holotype: MACEDONIA: distr . Kruševo, Kruševo env., beechwood 1 km NW of city, 1340 m (41°22'36.94"N, 21°14'8.74"E), 25.V.2014, ♂, J. Mertlik leg. ( PCJM) . Paratypes: MACEDONIA: same data as for the holotype, 42 ♂♂, 10 ♀♀, J. Mertlik leg. ( PCJM), dtto, 2 ♂♂, 1 ♀ ( PCRK) ; 32 ♂♂, 9 ♀♀, B. Zbuzek leg. (PCBZ); dtto, 34 ♂♂, 12 ♀♀, P. Brůha leg. (PCPB); Pelagonia reg., Krusevo, Fagus forest, singled at night, 41°22'38.01"N, 21°14'0.25"E, 5.VI.2014, 8 ♂♂, 4 ♀♀, leg. V. Dušánek, R. Kundrata & T. Németh ( HNHM) ; dtto, 39 ♂♂, 25 ♀♀ (PCRK); dtto, 17 ♂♂, 17 ♀♀, V. Dušánek leg. (PCVD); prov. Krusevo, Krusevo, Crn Viv , 1350 m, 1997. V.5.–6., 1 ♂, leg. A. Podlussány & I. Rozner ( HNHM) .

Other material examined. MACEDONIA: prov. Skopje, Skopska Crna Gora , 500–600 m , Mon. Sv. Jovan, 8.VI.1998, 2 ♀♀, leg. I. Rozner ( HNHM). These specimens were probably mislabelled (I. Rozner, pers. comm.).

Diagnosis. Dima dusaneki sp. nov. is a medium-sized to large species with more convex, suboval body and shiny elytral intervals with very sparsely distributed thin setae ( Figs 2 0–21). It shares the similar body coloration, pronotal punctation and the pubescence of the pronotal sides with D. raineri (shorter decumbent at anterior 1/2 or 2/3, longer semi-erect posteriorly; Figs 123 , 154–155 ). However, D. dusaneki sp. nov. differs from D. raineri (and also other Dima species in the region) in having a robust body and wide, laterally evenly rounded pronotum, which is widest medially ( Figs 20–21 ). Dima macedonica Schimmel, 1993 and D. florinensis differ from D. dusaneki sp. nov. in the more distinctly double-sized punctation of pronotum and the pubescence of the pronotal sides which is semi-erect along the whole margin length (shorter anteriorly in D. macedonica , long in D. florinensis ; Figs. 129 , 141–143 ). Dima fialai sp. nov. has the pronotum widest at posterior half, narrowed toward anterior margin, with sparser punctation ( Figs 29–30 ), and the pubescence of the pronotal sides short and almost decumbent at anterior 3/ 4 of the margin length and long semi-erect setae only near hind angles ( Fig. 128 ).

Description. Holotype, male. Body medium-sized, 12.5 mm long, 5.2 mm wide, shiny, covered with short dense pubescence. Body blackish-brown, with paler, reddish-brown legs, antennae, elytral sutura and margins; pubescence yellowish ( Fig. 20 ).

Head including eyes 0.5 times as wide as pronotum, frons slightly depressed; punctation sparse and coarse, moderately deep, intervals between punctures shiny; pubescence on head semi-erect, more erect in anterior part of frons, directed forwards. Supraantennal carina short, indistinct. Antennae moderately long, surpassing the posterior angles of pronotum of about 3.5 segments; antennomeres II and III subtriangular, antennomere II longer than antennomere III, both less than 1.5 times longer than wide, together 1.3 times longer than antennomere IV, length ratio of antennomeres II–IV 1.2: 1.0: 1.7, antennomeres IV to X gradually widened apically, about 2 times longer than wide, apical antennomere longest, subacute apically; surface of antennomeres covered with moderately long, semi-erect setae.

Pronotum 1.4 times wider than long, widest just behind middle, convex dorsally; sides convex; gradually narrowed towards frontal margin; anterior angles obtuse; posterior angles prominent, sharp, slightly produced outward; punctation sparse, moderately fine, punctures on disc equally distributed, of double size; intervals shiny. Pubescence on pronotal disc short, decumbent; that on pronotal sides at anterior 1/2 or 3/5 short and decumbent, pubescence longer and semi-erect to erect posteriorly, long and erect at hind angles ( Fig. 123 ). Prosternum finely, sparsely punctate, with short semi-erect setae, sparse medially, denser at margins; prosternal lobe moderately coarsely punctate, with longer semi-erect pubescence.

Scutellum sub-pentagonal, with sides rounded, with shallow median depression, in lateral view slightly raised above the plane of elytra, widest at middle; frontal margin rounded, gradually declined; apex widely rounded, punctation sparse, coarse; pubescence fine, moderately long, semi-erect. Elytra sub-parallel, 3.3 times longer than pronotum, convex, widest at posterior two thirds; striae well developed along the whole elytral length, irregularly interrupted by coarse sparse punctures; surface of interstices flattened, shiny, densely and finely punctate; punctures suboval, equally distributed, of the same size; pubescence short, sparse.

Aedeagus elongate; paramera long, narrow, its apical lobe moderately long, subapical tooth subacute ( Fig. 181 ).

Female ( Fig. 21 ). Like male, but more robust, with shorter antennae (surpassing the posterior angles of pronotum about one segment), pronotum distinctly wider than in male (1.6 times wider than long), scutellum distinctly wider. The sclerotized spines of bursa copulatrix as in Fig. 224 .

Intraspecific variability. Body length: 10.6–15.2 mm.

Distribution. Macedonia (Kruševo; Fig. 244 ).

Etymology. The specific epithet is a patronym in honor of Mr. Václav Dušánek (Zábřeh, Czech Republic), who collected a part of the type series.

Figures

FIGURE 244. The distribution of Dima species in Albania, Bulgaria, Macedonia and in the northern Greece.

FIGURES 15 – 29. Habitus images of Dima species. Dima dalmatina Küster, 1844, 15 — syntype, male, 16 — non-type male from Vosac, Croatia, 17 — non-type female from Vosac, Croatia, 18 — non-type male from Rumija Mts., Montenegro, 19 — nontype female from Rumija Mts., Montenegro; D. dusaneki sp. nov., 20 — holotype, male, 21 — paratype, female; D. elateroides Charpentier, 1825, 22 — non-type male from Serbia; 23 — non-type male from Slovenia, 24 — non-type female from Croatia; 25 — D. etoliensis Platia, 2012, holotype, female; D. evritaniensis Schimmel & Platia, 2008, 26 — paratype, female, Panetoliko Mts., 27 — non-type male from Timfristos Mts., 28 — non-type female from Timfristos Mts.; 29 — D. fialai sp. nov., holotype, male. Not to scale.

FIGURES 119 – 130. Pronotal details of Dima species. 119 — D. arndti Platia, 2013, male; 120 — D. assingi Schimmel & Platia, 2008, female; 121 — D. bruhai sp. nov., holotype, male; 122 — D. dalmatina Küster, 1844, male; 123 — D. dusaneki sp. nov., holotype, male; 124 — D. elateroides Charpentier, 1825, male; 125 — D. etoliensis Platia, 2012, holotype, female; 126 — D. evritaniensis Schimmel & Platia, 2008, paratype, female; 127 — D. evritaniensis, male; 128 — D. fialai sp. nov., holotype, male; 129 — D. florinensis Platia, 2012, female; 130 — D. fthiotidensis Schimmel & Platia, 2008, male. Not to scale.

FIGURES 172 – 181. Aedeagus images of Dima species. 172 — D. arndti Platia, 2013, holotype; 173 — D. arndti, non-type specimen; 174 — Dima sp. from Crete; 175 — D. assingi Schimmel & Platia, 2008, holotype; 176 — D. assingi, non-type specimen from Kilini Mts.; 177 — D. peloponnensis Schimmel & Platia 2008, holotype; 178 — D. bruhai sp. nov., holotype; 179 — D. dalmatina Küster, 1844, non-type specimen from Sutorman, Montenegro; 180 — D. dalmatina, non-type specimen from Vosec, Croatia; 181 — D. dusaneki sp. nov., holotype. Not to scale.

FIGURES 222 – 233. The sclerotized spines of bursa copulatrix of Dima species. 222 — D. arndti Platia, 2013, paratype; 223 — D. bruhai sp. nov., paratype; 224 — D. dusaneki sp. nov., paratype; 225 — D. elateroides, Charpentier, 1825, non-type specimen; 226 — D. etoliensis Platia, 2012, holotype; 227 — D. fialai sp. nov., paratype; 228 — D. florinensis Platia, 2012, holotype; 229 — D. fthiotidensis Schimmel & Platia, 2008, holotype of D. scutellaris Platia, 2010; 230 — D. giachinoi Schimmel & Platia 2008, paratype; 231 — D. hirtipennis Platia, 2011, holotype; 232 — D. kozufensis sp. nov., paratype; 233 — D. neumanni Platia, 2013, non-type specimen. Not to scale.

FIGURES 1 – 14. Habitus images of Dima species. Dima arndti Platia, 2013, 1 — holotype, male, 2 — paratype, female, 3 — nontype specimen, male, 4 — non-type specimen, female; 5 — Dima sp. from Crete, male; D. assingi Schimmel & Platia, 2008, 6 — holotype, male, 7 — holotype of D. peloponnensis Schimmel & Platia, 2008, male, 8 — non-type male from Kilini Mts., 9 — nontype female from Kilini Mts.; D. bruhai sp. nov., 10 — holotype, male, 11 — paratype, female, 12 — paratype, male, 13 — paratype, male; D. dalmatina Küster, 1844, 14 — syntype, male, lateral view. Not to scale.

FIGURES 143 – 154. Pronotal details of Dima species. 143 — D. macedonica Schimmel, 1993, male; 144 — D. marvani Mertlik & Dusanek, 2006, holotype, male; 145 — D. moraveci Schimmel & Platia, 2008, holotype, female; 146 — D. neumanni Platia, 2013, female; 147 — D. olympica Meschnigg, 1934, male; 148 — D. orientalis sp. nov., holotype, male; 149 — D. parnonensis Schimmel & Platia, 2008, female; 150 — D. pecoudi Fleutiaux, 1944, male; 151 — D. pelikani sp. nov., holotype, male; 152 — D. pelionensis sp. nov., paratype, female, 153 — D. pelionensis sp. nov., paratype, female paratype of D. fokidensis; 154 — D. raineri Wurst, 1997, female. Not to scale.

FIGURES 155 – 162. Pronotal details of Dima species. 155 — D. raineri Wurst, 1997, holotype, male; 156 — D. schimmeli Platia & Gudenzi, 2009, female; 157 — D. schnitteri Platia, 2013, female; 158 — D. vailatii Schimmel & Platia 2008, holotype, male; 159 — D. vailatii, male; 160 — D. riesei Platia, 2010, holotype, male; 161 — D. vonickai sp. nov., holotype, male; 162 — D. zbuzeki sp. nov., holotype, male. Not to scale.

FIGURES 131 – 142. Pronotal details of Dima species. 131 — D. fthiotidensis Schimmel & Platia, 2008, paratype, female; 132 — D. scutellaris Platia, 2010, holotype, female; 133 — D. fthiotidensis, male; 134 — D. fthiotidensis, holotype of D. fokidensis Schimmel & Platia, 2008; 135 — D. giachinoi Schimmel & Platia 2008, female; 136 — D. hirtipennis Platia, 2011, holotype, female; 137 — D. hladilorum Schimmel, 1987, male; 138 — D. kozufensis sp. nov., paratype, female; 139 — Dima lebenbaueri Schimmel & Platia 2008, holotype, male; 140 — Dima lebenbaueri, non-type specimen, female; 141 — D. macedonica Schimmel, 1993, holotype, female, 142 — D. macedonica, female paratype of D. hladilorum from Galičica Mts. Not to scale.