Dima hladilorum Schimmel, 1987

Mertlik, Josef, Németh, Tamás & Kundrata, Robin, 2017, Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula, Zootaxa 4220 (1), pp. 1-63 : 19

Dima hladilorum Schimmel, 1987

Figs 47–49 , 137 , 190 , 245 .

Dima hladilorum Schimmel, 1987: 42 ; Schimmel & Platia 1991: 293; Schimmel 1993: 247; Schimmel 1996: 141; Wurst 1997: 3; Cate 2007: 184; Schimmel & Platia 2008: 580; Platia 2012: 200.

Type depositories. Holotype, ♂ (PCRS), allotype, ♀ (PCJH), 4 paratypes: 2 ♂♂, 2 ♀♀ (PCRS, PCGP, PCJM, PCJH).

Type locality. Greece, Parnassos Mt., 1700 m.

Redetermined material. Three paratypes of D. hladilorum from Macedonia (Galičica Mts.; Schimmel 1987) belong to D. macedonica (described later than D. hladilorum ; Schimmel 1993) ( Fig. 61 ). Some specimens recorded under D. hladilorum in Schimmel & Platia (2008) were redetermined as D. arndti (Taygetos Mts., Crete; 4 ex.), D. assingi (Kilini Mts.; 1 ex.), D. pelionensis sp. nov. (Mt. Pilio; 2 ex.), and D. vailatii (Kajmakčalan Mts.; 2 ex.).

New material. Greece: distr. Viotía , Parnassos Mts., Ski center env., firrywood, 1700 m (38°32'54.89"N, 22°34'25.84"E), 28.V.2013, 9 ♂♂, 10 ♀♀, J. Mertlik leg. ( PCJM) ; dtto, 1 ♂, 2 ♀♀ (PCRK); dtto, 6 ♂♂, 8 ♀♀, V. Dušánek leg. (PCVD); dtto, 6 ♂♂, 8 ♀♀, P. Brůha leg. (PCPB); distr. Viotía , Parnassos Mts., Ski center env., firrywood, 1700 m (38°32'54.89"N, 22°34'25.84"E), 30.V.2014, 41 ♂♂, 15 ♀♀, J. Mertlik leg. ( PCJM) ; dtto, 32 ♂♂, 10 ♀♀, B. Zbuzek leg. (PCBZ); dtto, 12 ♂♂, 14 ♀♀, P. Brůha leg. (PCPB); prov. Central Gr. , distr. Phthiotis, Mt. Parnassos, 4 km NE Kalivia Arachova, spruce forest, 38°32'22.65"N, 22°34'12.45"E, 1700 m, singled at night, 25.–26.V.2015, leg. A. Kotán, Gy. Németh, B. Szelenczey & V. Szénási, 9 ♂♂, 6 ♀♀ ( PCBS) ; dtto, 10 ♂♂, 3 ♀♀ (PCRK); dtto, 2 ♂♂, 2 ♀♀ (HNHM).

Diagnosis. Dima hladilorum is a smaller species (body length: 9.4–10.0 mm; Figs 47–49 ) with pronotum with sparse punctuation and shiny intervals, pubescence of the pronotal sides semi-erect along its whole length ( Fig. 137 ), scutellum convex with its frontal margin steeply declined, and paramera robust, elongate ( Fig. 190 ). This species is like D. zbuzeki sp. nov. from Agrafa Mts., however it differs from the latter by the less distinct pubescence on the posterior angles of pronotum, convex scutellum with medially emarginate frontal margin, elytral striae often indistinct or missing in the apical part, and by the shape of the apical lobe of paramera ( Figs 47-49 , 116–118 , 137 , 162 , 190 , 221 ). Dima fthiotidensis and D. evritaniensis differ from D. hladilorum in the matt, densely punctate pronotum ( Figs 26–28 , 35–41 ) and smaller apical lobe of paramera ( Figs 183, 186–187 ); D. etoliensis by the shape of antennae (narrower, with longer antennomeres), flattened scutellum with gradually declined frontal margin, larger body with more parallel-sided elytra, and the distinct striae along the whole length of elytra ( Fig. 25 ). Dima pelionensis sp. nov. has a flattened scutellum, pubescence of the pronotal sides decumbent in apical two thirds ( Figs 152–153 ), and the apical lobe of paramera shorter ( Fig. 209 ).

Intraspecific variability. Some specimens have more or less indistinct elytral striae.

Distribution. Greece (Parnassos Mts.; Fig. 245 ).

Figures

FIGURE 245. The distribution of Dima species in the Central Greece.

FIGURES 45 – 59. Habitus images of Dima species. 45 — Dima hirtipennis Platia, 2011, holotype, female, 46 — Dima sp. from Mani; D. hladilorum Schimmel, 1987, 47 — paratype, female, 48 — non-type specimen, male, 49 — non-type specimen, female; D. kozufensis sp. nov., 50 — holotype, male, 51 — paratype, female, 52 — paratype, female, paratype of D. lebenbaueri Schimmel & Platia, 2008; D. lebenbaueri Schimmel & Platia 2008, 53 — holotype, male, 54 — paratype, female, 55 — non-type specimen, male, 56 — non-type specimen, female; D. macedonica Schimmel, 1993, 57 — holotype, male, 58 — non-type male from Galičica, 59 — non-type female from Galičica Mts. Not to scale.

FIGURES 131 – 142. Pronotal details of Dima species. 131 — D. fthiotidensis Schimmel & Platia, 2008, paratype, female; 132 — D. scutellaris Platia, 2010, holotype, female; 133 — D. fthiotidensis, male; 134 — D. fthiotidensis, holotype of D. fokidensis Schimmel & Platia, 2008; 135 — D. giachinoi Schimmel & Platia 2008, female; 136 — D. hirtipennis Platia, 2011, holotype, female; 137 — D. hladilorum Schimmel, 1987, male; 138 — D. kozufensis sp. nov., paratype, female; 139 — Dima lebenbaueri Schimmel & Platia 2008, holotype, male; 140 — Dima lebenbaueri, non-type specimen, female; 141 — D. macedonica Schimmel, 1993, holotype, female, 142 — D. macedonica, female paratype of D. hladilorum from Galičica Mts. Not to scale.

FIGURES 182 – 191. Aedeagus images of Dima species. 182 — D. elateroides Charpentier, 1825, non-type specimen from Serbia; 183 — D. evritaniensis Schimmel & Platia, 2008, non-type specimen from Timfristos Mts.; 184 — D. fialai sp. nov., holotype; 185 — D. florinensis Platia, 2012, non-type specimen; 186 — D. fthiotidensis Schimmel & Platia, 2008, non-type specimen from Kallidromo Mts.; 187 — D. fthiotidensis, non-type specimen from Vardoussia Mts.; 188 — D. giachinoi Schimmel & Platia 2008, non-type specimen; 189 — Dima sp. from Mani; 190 — D. hladilorum Schimmel, 1987, non-type specimen; 191 — D. kozufensis, sp. nov., holotype. Not to scale.

FIGURES 60 – 74. Habitus images of Dima species. Dima macedonica Schimmel, 1993, 60 — non-type male from Ehloec, 61 — female paratype of D. hladilorum Schimmel, 1987 from Galičica Mts.; D. marvani Mertlik & Dusanek, 2006, 62 — holotype, male, 63 — female paratype from Lovčen Mts., 64 — female paratype from Rumija Mts., 65 — non-type male from Rumija Mts.; 66 — D. moraveci Schimmel & Platia 2008, holotype, female; D. neumanni Platia, 2013, 67 — holotype, male, 68 — non-type male from Erimanthos Mts., 69 — non-type female from Erimanthos Mts., 70 — non-type male from Aroania Mts., 71 — non-type female from Aroania Mts., 72 — non-type female from Aroania Mts., 73 — non-type female from Panachaiko Mts., 74 — non-type female from Panachaiko Mts. Not to scale.

FIGURES 105 – 118. Habitus images of Dima species. Dima schnitteri Platia, 2013, 105 — holotype, male, 106 — paratype, female, 107 — non-type specimen, male, 108 — non-type specimen, female; D. vailatii Schimmel & Platia 2008, 109 — holotype, male, 110 — paratype, female, 111 — non-type specimen, male, 112 — non-type specimen, female, 113 — holotype of D. riesei Platia, 2010, male; D. vonickai sp. nov., 114 — holotype, male, 115 — paratype, female; D. zbuzeki sp. nov., 116 — holotype, male, 117 — paratype, female, 118 — paratype, male. Not to scale.

FIGURES 155 – 162. Pronotal details of Dima species. 155 — D. raineri Wurst, 1997, holotype, male; 156 — D. schimmeli Platia & Gudenzi, 2009, female; 157 — D. schnitteri Platia, 2013, female; 158 — D. vailatii Schimmel & Platia 2008, holotype, male; 159 — D. vailatii, male; 160 — D. riesei Platia, 2010, holotype, male; 161 — D. vonickai sp. nov., holotype, male; 162 — D. zbuzeki sp. nov., holotype, male. Not to scale.

FIGURES 212 – 221. Aedeagus images of Dima species. 212 — D. raineri Wurst, 1997, paratype of D. macedonica; 213 — D. raineri, holotype of D. pindosensis Schimmel & Platia, 2008; 214 — D. schimmeli Platia & Gudenzi, 2009, holotype; 215 — D. schimmeli, non-type specimen; 216 — D. schnitteri Platia, 2013, holotype; 217 — D. vailatii Schimmel & Platia, 2008, holotype; 218 — D. vailatii, non-type specimen; 219 — D. vailatii, holotype of D. riesei Platia, 2010; 220 — D. vonickai sp. nov., holotype; 221 — D. zbuzeki sp. nov., holotype. Not to scale.

FIGURES 15 – 29. Habitus images of Dima species. Dima dalmatina Küster, 1844, 15 — syntype, male, 16 — non-type male from Vosac, Croatia, 17 — non-type female from Vosac, Croatia, 18 — non-type male from Rumija Mts., Montenegro, 19 — nontype female from Rumija Mts., Montenegro; D. dusaneki sp. nov., 20 — holotype, male, 21 — paratype, female; D. elateroides Charpentier, 1825, 22 — non-type male from Serbia; 23 — non-type male from Slovenia, 24 — non-type female from Croatia; 25 — D. etoliensis Platia, 2012, holotype, female; D. evritaniensis Schimmel & Platia, 2008, 26 — paratype, female, Panetoliko Mts., 27 — non-type male from Timfristos Mts., 28 — non-type female from Timfristos Mts.; 29 — D. fialai sp. nov., holotype, male. Not to scale.

FIGURES 30 – 44. Habitus images of Dima species. 30 — D. fialai sp. nov., paratype, female; D. florinensis Platia, 2012, 31 — holotype, female, 32 — non-type male from Varnous Mts., 33 — non-type female from Varnous Mts., 34 — non-type female from Galičica Mts.; D. fthiotidensis Schimmel & Platia, 2008, 35 — paratype, female, 36 — holotype of D. fokidensis Schimmel & Platia, 2008, female, 37 — holotype of D. scutellaris Platia, 2010, female, 38 — non-type male from Kallidromo Mts., 39 — nontype male from Vardoussia Mts., 40 — non-type female from Vardoussia Mts., 41 — non-type female from Vardoussia Mts.; D. giachinoi Schimmel & Platia 2008, 42 — paratype, female, 43 — non-type specimen, male, 44 — non-type specimen, female. Not to scale.

FIGURES 143 – 154. Pronotal details of Dima species. 143 — D. macedonica Schimmel, 1993, male; 144 — D. marvani Mertlik & Dusanek, 2006, holotype, male; 145 — D. moraveci Schimmel & Platia, 2008, holotype, female; 146 — D. neumanni Platia, 2013, female; 147 — D. olympica Meschnigg, 1934, male; 148 — D. orientalis sp. nov., holotype, male; 149 — D. parnonensis Schimmel & Platia, 2008, female; 150 — D. pecoudi Fleutiaux, 1944, male; 151 — D. pelikani sp. nov., holotype, male; 152 — D. pelionensis sp. nov., paratype, female, 153 — D. pelionensis sp. nov., paratype, female paratype of D. fokidensis; 154 — D. raineri Wurst, 1997, female. Not to scale.

FIGURES 202 – 211. Aedeagus images of Dima species. 202 — D. olympica Meschnigg, 1934, non-type specimen; 203 — D. orientalis sp. nov., holotype; 204 — D. parnonensis Schimmel & Platia 2008, non-type specimen; 205 — D. parnonensis, nontype specimen; 206 — D. pecoudi Fleutiaux, 1944, paratype of D. evritaniensis; 207 — D. pecoudi, non-type specimen; 208 — D. pelikani sp. nov., holotype; 209 — D. pelionensis sp. nov., paratype; 210 — D. raineri Wurst, 1997, holotype; 211 — D. raineri, non-type specimen. Not to scale.