Fannia amplitarsis, Gomes & Pedraza-Lara, 2023

Gomes, Lucas R. P. & Pedraza-Lara, Carlos, 2023, Fannia pretiosa-group (Diptera: Fanniidae) from Mexico: six new species from the Mexico Transition Zone and an identification key, Zootaxa 5264 (2), pp. 185-204 : 189-190

publication ID	https://doi.org/ 10.11646/zootaxa.5264.2.2
publication LSID	lsid:zoobank.org:pub:97A03469-519D-49D7-8D0E-2F6648CCB922
DOI	https://doi.org/10.5281/zenodo.7836359
persistent identifier	https://treatment.plazi.org/id/0396A100-C21A-EE76-DDF7-FBD643611484
treatment provided by	Plazi
scientific name	Fannia amplitarsis
status	sp. nov.

Treatment

Fannia amplitarsis View in CoL sp. nov.

( Figs 1A, 1C View FIGURE 1 , 3B View FIGURE 3 , 4B View FIGURE 4 , 5B View FIGURE 5 , 6B View FIGURE 6 , 7 View FIGURE 7 , 16 View FIGURE 16 )

Diagnosis. Fore tarsi white-yellowish, fore tarsomeres 1–4 strongly expanded and flattened, tarsomere 5 bearing 1 ad long setula (three times longer than tarsomere) ( Fig. 7D View FIGURE 7 ). Hind coxa with two long setulae posteriorly ( Fig. 1C View FIGURE 1 ). Hind femur slightly curved with pre-apical swelling on v surface, with a tuft of long setae (three times longer than width of femur) on the swelling, including two sets of 3–4 long setae, one on av surface and another on the pv surface (the pv setae slightly longer, thicker and apically curled) ( Fig. 7F View FIGURE 7 ). Wing brownish ( Fig. 7C View FIGURE 7 ).

Description. Holotype male. Body length: 6.8 mm. Wing length: 5.8 mm.

Head. Eye densely setulose. Ocellar triangle black; ocellar setae proclinate and developed (as long as frontal setae). Frontal vitta brownish-red, at the narrowest point three times wider than width of anterior ocellus ( Fig. 7A View FIGURE 7 ). Frontal setae 17. Fronto-orbital plate brown with silver pruinosity. Parafacial bare, brownish-red with silver pruinosity. Frontogenal suture and gena silver pruinose. Gena brownish-red with silver pruinosity ( Fig. 7A View FIGURE 7 ). Pedicel with dorsal seta developed, almost reaching half-length of arista. Postpedicel black, golden pilose.

Thorax. Scutum without vittae, inconspicuously silver pruinose on postpronotal and notopleural regions ( Fig. 7B View FIGURE 7 ). Acrostichal setulae triserial presuturally and quadriserial postsuturally. Dorsocentrals 2+3, long, the posteriormost postsutural pair 1/3 longer than the others. Prealar 2, the anterior slightly longer and thicker than posterior one. Katepisternum with about 15 long and thin setae ventrally ( Fig. 7C View FIGURE 7 ).

Wing. Hyaline ( Figs 7B, 7C View FIGURE 7 ). Calypters white. Haltere yellow, base of stem brown.

Legs. Black, pulvilli brownish and tarsi brown, except for the fore tarsomeres 1–4 which are white-yellowish. Fore tarsomeres enlarged, first tarsomere enlarged apically and with 2 v setulae, tarsomere 2 enlarged, longer than its width, tarsomere 3–5 enlarged, wider than its length, tarsomeres 2–4 brown and yellow, tarsomere 5 yellow, with long setae, four times longer than tarsomere 5 width ( Fig. 7D View FIGURE 7 ). Fore tarsomere 2 with a ventral spike apically. Mid femur with 1 row of ad setae; with 1 row of decreasing av from base to apex; 2 rows of long pv, a tuft of short v setae on preapical third ( Fig. 7E View FIGURE 7 ). Mid tibia with 2 submedian ad, 1 apical setae on pd, p, and pv, and 2 av, and 1 long a apical setae twice longer than the others ( Fig. 7E View FIGURE 7 ). Hind coxa with 2 setae on posterior margin ( Fig. 1C View FIGURE 1 ). Hind femur curved with pre-apical swelling on v surface, with a tuft of long setae on the swelling, including a set of 3 long av setae and 4 long pv setae (the pv setae slightly longer, thicker and apically curled), 1 row of long (twice longer than width of femur) ad to d setae (with several setae between these rows) from base to mid-third, a row of pv setae, 1 long preapical av setae ( Fig. 7F View FIGURE 7 ). Hind tibia with 4 median av setae; 1 apical av; a and ad surfaces covered with a full row of short setae and 1 short a and av apical setae ( Fig. 7F View FIGURE 7 ).

Abdomen. Metallic black-bluish with silver pruinosity, elongate, with developed setae,tergite 5 with long marginal setae (slightly longer than others). Black pruinosity forms a central longitudinal stripe dorsally. Syntergosternite with 2 short lateral setae. Sternite 5 quadrangular, posterior margin profoundly indented ( Fig. 3B View FIGURE 3 ).

Terminalia. Surstyli fused with epandrium. Surstyli slightly curved apically, with a medial projection outward directed ( Figs 4B View FIGURE 4 , 5B View FIGURE 5 ). Bacilliform process long and simple, as long as half of the surstyli ( Fig. 4B View FIGURE 4 ). Hypandrial arms directed outwards ( Fig. 6B View FIGURE 6 ).

Female. Unknown.

Etymology. The specific name is derived from the Latin words ampli - (“wide”) and tarsis (“tarsi”), which refers to the modified fore tarsi of male.

Type-material. Holotype. Male. Mexico. Tlaxcala. La Malinche [19.24432, -98.09781], 3480 m, 29.vii.1989, E. Ramírez ( CNIN) GoogleMaps . Paratypes. Mexico. Mexico. Tequesquinahuac, C. Tlaloc [19.3987, -98.7085], 15.iv.1982, J. Butze, 2 males ( CNIN) GoogleMaps .

Distribution. Mexico (State of Mexico, Tlaxcala).

Remarks. The fore tarsi of F. amplitarsis is enlarged and similar to the Chilean species F. schnusei Stein, 1911 ( Domínguez 2007) , but differ in the shape of tarsomeres and absence of an apical thorn on the posterior margin. It also differs in the mid tarsomere 1, by the absence of a strong ventral basal crest; mid femur not strongly curved, without a strong preapical protuberance on av surface; hind coxae setulose posteriorly; legs chaetotaxy; and male terminalia, mainly in the surstyli and process bacilliform shape.