Eckhartia longicolumella E.M.FRIIS, P.R.CRANE et K.R.PEDERSEN, 2019

Friis, Else Marie, Crane, Peter R. & Pedersen, Kaj Raunsgaard, 2019, The Early Cretaceous Mesofossil Flora Of Torres Vedras (Ne Of Forte Da Forca), Portugal: A Palaeofloristic Analysis Of An Early Angiosperm Community, Fossil Imprint 75 (2), pp. 153-257 : 210-213

publication ID

https://doi.org/ 10.2478/if-2019-0013

persistent identifier

https://treatment.plazi.org/id/0396DC10-BF36-C23F-CE32-B5B8E7961FBE

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Diego

scientific name

Eckhartia longicolumella E.M.FRIIS, P.R.CRANE et K.R.PEDERSEN
status

sp. nov.

Eckhartia longicolumella E.M.FRIIS, P.R.CRANE et K.R.PEDERSEN sp. nov.

Text-fig. 38a–i View Text-fig

H o l o t y p e. Designated here. S136754 (Torres Vedras sample 44; figured Text-fig. 38b–i View Text-fig ).

P l a n t F o s s i l N a m e s R e g i s t r y N u m b e r.

PFN000473 (for new species).

P a r a t y p e s. Designated here. S136673, S137901

(Torre Vedras sample 44).

R e p o s i t o r y. Palaeobotanical Collections , Department of Palaeobiology, the Swedish Museum of Natural History, Stockholm, Sweden .

E t y m o l o g y. From Latin:longus for long and columella referring to the long columellae supporting the muri.

T y p e l o c a l i t y. Torres Vedras (NE of Forte de

Forca; 39°06′13″ N, 9°14′47″ W).

T y p e s t r a t u m a n d a g e. Lower member of the Almargem Formation; Early Cretaceous (late Barremianearly Aptian).

D i a g n o s i s. As for the genus with the following additions: Anthers long and narrow. Reticulum fine. Columellae long, scattered.

D i s t i n g u i s h i n g f e a t u r e s. See comments for

Eckhartia brevicolumella and E. intermedia .

D i m e n s i o n s. Length of anthers: about 0.55 mm; width of anthers: 0.15 mm. Length of pollen grains: 16–19 µm.

D e s c r i p t i o n a n d r e m a r k s. Eckhartia longicolumella is based on a single well-preserved anther and several anther fragments. It was first illustrated by Friis et al. (2010a), but not formally described or named. The anthers are about 0.55 mm long, 0.15 mm wide and are dithecate and tetrasporangiate ( Text-fig. 38a, b View Text-fig ).

Pollen grains are about 16–19 µm in equatorial diameter and monocolpate. The exine is semitectate-reticulate, heterobrochate and columellate ( Text-fig. 38c–i View Text-fig ). The reticulum over most of the pollen grain is relatively fine compared to the reticulum of Eckhartia brevicolumella . Lumina are of two sizes; larger lumina are up to about 1.35 µm in diameter ( Text-fig. 38g View Text-fig ). Towards the colpus margin, and particularly around the end of the colpi, the lumina become very small and the colpus margin is psilate ( Text-fig. 38c, e, f View Text-fig ). Muri are about 0.2 µm wide with a rounded or flattened profile. Columellae are long, about 0.45 µm long, typically widely spaced ( Text-fig. 38g, i View Text-fig ). The reticulum is only loosely attached and easily separates from the foot layer, leaving the foot layer naked or with only columellae attached ( Text-fig. 38 h, i View Text-fig ). Orbicules are densely distributed on the inner surface of the anther wall and scattered among the pollen grains ( Text-fig. 38f View Text-fig ). The orbicules are closely similar to the tooth-like orbicules of E. brevicolumella ( Text-fig. 38f View Text-fig ).

A f f i n i t y a n d o t h e r o c c u r r e n c e s. Like Eckhartia brevicolumella , E. longicolumella has not been observed in any other mesofossil flora from Portugal. The grains of E. longicolumella resemble the grains found on the stigmatic surface of Canrightia sp. described here (p. 183 herein), but differ in the shape of the muri and details of the reticulum.

R

Departamento de Geologia, Universidad de Chile

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