Athetina Casey, 1910a

Asenjo, Angélico, Irmler, Ulrich, Klimaszewski, Jan, Chandler, Donald S., Fierros-López, Hugo Eduardo & Vieira, Juliana S., 2019, Staphylinidae (Insecta: Coleoptera) in Latin America: synopsis, annotated catalog, diversity and distribution, Zootaxa 4621 (1), pp. 1-406 : 93-103

Subtribe Athetina Casey, 1910a

0300. Acrotona Thomson, 1859: 38

Type species: Aleochara aterrima Gravenhorst, 1802

Eustrigota Casey, 1911: 165 [synonymized by Gusarov 2 003e: 102]

Type species: Strigota seclusa Casey, 1911

Pancota Casey, 1906: 345 [ Newton & Thayer 2003]

Type species: Pancota collaris Casey, 1906

Subgenera:

Acrotona Thomson, 1859: 38

Type species: Aleochara aterrima Gravenhorst, 1802

Colpodota Mulsant & Rey, 1873d: 153 [synonymized by Schülke & Smetana 2015: 506]

Type species: Homalota parens Mulsant & Rey, 1852

Colpodota Mulsant & Rey, 1873b: 175 [ Newton & Thayer 2003]

Colpodota Mulsant & Rey, 1874c: 207 [ Newton & Thayer 2003]

Colpodota Mulsant & Rey, 1874a: 7 [ Newton & Thayer 2003]

Aremia Casey, 1910a: 145

Type species: Dolosota reclusa Casey, 1910

Arisota Casey, 1910a: 133

Type species: Arisota tetricula Casey, 1910

Dolosota Casey, 1910a: 136

Type species: Eurypronota scopula Casey, 1893

Eurypronota Casey, 1893: 334

Type species: Eurypronota discreta Casey, 1893

Neada Casey, 1910a: 152

Type species: Acrotona lubricans Casey, 1910

Phanerosphena Pace, 1991a: 138

Type species: Atheta rougemonti Pace, 1991

Reania Casey, 1910a: 146

Type species: Dolosota fontinalis Casey, 1910

56 species are included in this genus from Latin America.

Characters/Description: Lynch-Arribálzaga 1884: 58 (characters of Colpodota ); Fenyes 1920: 202 (characters as Pancota ), 223 (characters); Lohse et al. 1990: 126 (characters in key), 198 (characters).

Key: Newton & Peck 2006: 139 (species of the Galapagos Isl.).

Distribution.

West Indies: Jamaica, Haiti, Guadeloupe, Saint Vincent & the Grenadines, Grenada.

Central America: Guatemala, Costa Rica.

South America: Colombia, French Guiana, Ecuador, Galapagos Isl., Peru, Bolivia, Brazil, Paraguay, Chile, Argentina.

Other regions: Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian, Oceanic.

0301. Adota Casey, 1910a: 67

Type species: Atheta massettensis Casey, 1910

Halostiba Yosii & Sawada, 1976: 86 [synonymized by Gusarov 2003b: 4]

Type species: Ischnopoda ushio Sawada, 1971

Panalota Casey, 1910a: 71 [ Lohse & Smetana 1985: 282]

Type species: Atheta setositarsis Casey, 1910

Phyconoma Easton, 1971: 24 [synonymized by Lohse 1988: 46]

Type species: Atheta immigrans Easton, 1971

1 species is included in this genus from Central America.

Characters/Description: Fenyes 1920: 175 (characters), 243 (characters as Panalota ); Gusarov 2003b: 3 (characters).

Key: Gusarov 2003b: 8 (Nearctic species).

Distribution.

Central America: Mexico.

Other regions: Nearctic, Palaearctic.

0302. Amidobia Thomson, 1858: 33

Type species: Homalota talpa Heer, 1841

1 species is included in this genus from South America.

Characters/Description: Fenyes 1920: 178 (characters).

Distribution.

South America: Brazil.

Other regions: Nearctic, Palaearctic, Oriental.

0303. Amischa Thomson, 1858: 33

Type species: Aleochara analis Gravenhorst, 1802

Arthropycna Bernhauer, 1921c: 162 [synonymized by Blackwelder 1944: 159]

Type species: Atheta myrmecovagans Bernhauer, 1921

Colposura Casey, 1893: 336 [synonymized by Bernhauer & Scheerpeltz 1926: 592]

Type species: Colposura praelonga Casey, 1893

15 species are included in this genus from Central and South America.

Characters/Description: Fenyes 1920: 178 (characters); Pace 2008a: 280 (characters in key).

Distribution.

Central America: Guatemala.

South America: Colombia, Ecuador, Brazil, Chile, Argentina.

Other regions: Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian, Oceanic.

0304. Atheta Thomson, 1858: 36 ( Figs. 48–49 )

Type species: Aleochara graminicola Gravenhorst, 1806

Iotota Casey, 1910a: 95 [ Newton & Thayer 2003]

Type species: Iotota tepida Casey, 1910

Megonychiella Marcuzzi, 1977: 17 [nomen nudum, Newton & Thayer 2003]

Type species: Atheta oxypodoides Marcuzzi, 1977

Paraceritaxa Marcuzzi, 1977: 16 [nomen nudum, Newton & Thayer 2003]

Type species: Atheta sexspinulosa Marcuzzi, 1977

Pseudometaxia Wüsthoff, 1940: 683 [ Blackwelder 1952: 328]

Rhodeota Casey, 1911: 147 [ Newton & Thayer 2003]

Type species: Ousipalia tartarea Casey, 1910

Synaptina Casey, 1910a: 131 [ Newton & Thayer 2003]

Type species: Synaptina merica Casey, 1910

Subgenera:

Acromyrmecoxene Scheerpeltz, 1976a: 138 [*]

Type species: Atheta acromyrmecicola Scheerpeltz, 1976

Acrotonida Pace, 1991a: 141

Type species: Atheta invicta Fauvel, 1907

Alaobia Thomson, 1858: 36

Type species: Homalota ochracea Erichson, 1837

Anopleta Mulsant & Rey, 1873b: 4 [*]

Type species: Homalota lepida Kraatz, 1856

Ouralia Mulsant & Rey, 1873d: 174 [ Smetana 2004: 372]

Type species: Ouralia picicornis Mulsant & Rey, 1873

Arctostiba Bernhauer, 1928b: 16

Type species: Atheta freyi Bernhauer, 1928

Atheta Thomson, 1858: 36 [*]

Type species: Aleochara graminicola Gravenhorst, 1806

Callicerodes Iablokoff-Khnzorian, 1960: 1883 [synonymized by Assing 2005: 63]

Type species: Callicerus velox Iablokoff-Khnzorian, 1960

Elytrusa Casey, 1906: 336 [ Smetana 2004: 372]

Type species: Homalota granulata Mannerheim, 1846

Hypatheta Fenyes, 1918: 23 [ Benick & Lohse 1974: 191]

Type species: Bolitochara castanoptera Mannerheim, 1830

Megista Mulsant & Rey, 1873b: 591 [ Benick & Lohse 1974: 191]

Type species: Aleochara graminicola Gravenhorst, 1806

Attatheta Scheerpeltz, 1936: 507 [*]

Type species: Atheta anisophthalma Scheerpeltz, 1936

Badura Mulsant & Rey, 1873d: 159

Type species: Badura nudicornis Mulsant & Rey, 1873

Bessobia Thomson, 1858: 35 [*]

Type species: Homalota monticola Thomson, 1852

Thrichiota Mulsant & Rey, 1873d: 180 [synonimized by Bernhauer & Scheerpeltz 1926: 625]

Type species: Bessobia gibbera Mulsant & Rey, 1873

Ceritaxa Mulsant & Rey, 1873d: 164

Type species: Ceritaxa spissata Mulsant & Rey, 1873

Hilara Mulsant & Rey, 1873: 160 [synonimized by Lohse 1971: 77]

Type species: Microdota fulva Mulsant & Rey, 1873

Neohilara Lohse, 1971: 77 [synonimized by Assing 2015a: 16]

Type species: Microdota fulva Mulsant & Rey, 1873

Chaetida Mulsant & Rey, 1873b: 272 [*]

Type species: Aleochara longicornis Gravenhorst, 1802

Coprothassa Thomson, 1859: 38 [*]

Type species: Homalota testudinea Erichson, 1839

Hemitropia Mulsant & Rey, 1873b: 179 [synonimized by Bernhauer & Scheerpeltz 1926: 670]

Type species: Oxypoda melanaria Mannerheim, 1830

Dabura Cameron, 1948a: 241

Type species: Atheta anommatophila Cameron, 1948

Datomicra Mulsant & Rey, 1873b: 355 [*]

Type species: Homalota celata Erichson, 1837

Datostiba Yosii & Sawada, 1976: 18 [ Silfverberg 1992: 29]

Type species: Atheta lewisiana Cameron, 1933

Hilarina Casey, 1910a: 128 [synonimized by Bernhauer & Scheerpeltz 1926: 666]

Type species: Datomicra particula Casey, 1910

Micromota Casey, 1910a: 127 [synonimized by Bernhauer & Scheerpeltz 1926: 666]

Type species: Datomicra filiformis Casey, 1910

Monadia Casey, 1910a: 130 [Synonimized by Bernhauer & Scheerpeltz 1926: 666]

Type species: Datomicra lucana Casey, 1910

Oligomia Casey, 1910a: 129 [synonimized by Bernhauer & Scheerpeltz 1926: 666]

Type species: Datomicra scintilla Casey, 1910

Taxicerella Casey, 1910a: 113 [ Smetana 2004: 379]

Type species: Sableta remissa Casey, 1910

Delphota Casey, 1910a: 17

Type species: Atheta cephalina Casey, 1910

Diaprepota Pace, 2004c: 360

Type species: Atheta ruiliensis Pace, 1998

Diaprepota Pace, 1998: 926 [synonimized by Pace 2004c: 360]

Type species: [no type species designated]

Dimetrota Mulsant & Rey, 1873d: 165 [*]

Type species: Homalota tristicula Mulsant & Rey, 1873

Cadaverota Yosii & Sawada, 1976: 55 [ Smetana 2004: 380]

Type species: Homalota cadaverina Brisout de Barneville, 1860

Coproceramius Gistel, 1857: 9 [ Smetana 2004: 380]

Type species: Bolitochara impressifrons Mannerheim, 1830

Engamota Casey, 1910a: 151 [ Smetana 2004: 380]

Type species: Acrotona absona Casey, 1910

Dimetrotina Casey, 1911: 143

Type species: Dimetrota vaniuscula Casey, 1911

Mycetota Ádám, 1987b: 13 [synonymized by Gusarov 2 016: 395]

Type species: Aleochara laticollis Stephens, 1832

Donesia Casey, 1910a: 48

Type species: Atheta temporalis Casey, 1910

Dysanomota Pace, 2008d: 431 [*]

Type species: Homalota cinctella Erichson, 1839

Ekkliatheta Pace, 2004b: 507

Type species: Atheta aniiensis Pace, 2004

Epimella Peyerimhoff, 1914: 250

Type species: Atheta cinctuta Peyerimhoff, 1914

Eubadura Sawada, 1990: 543

Type species: Atheta akiensis Sawada, 1990

Eugluta Sawada, 1985: 93

Type species: Atheta picticollis Bernhauer, 1915

Eumorphotheta Ádám, 2008: 161

Type species: Atheta siculorum Ádám, 2008

Evaniodera Pace, 1986a: 430 [*]

Type species: Atheta apicilutea Pace, 1986

Geostibops Israelson, 1985: 13

Type species: Atheta aptera Israelson, 1985

Gyroatheta Pace, 2014c: 100 [*]

Type species: Atheta guyanicola Pace, 2014

Hummleriella Bernhauer, 1929c: 191

Type species: Atheta ponferradae Bernhauer, 1929

Hydrosmectomorpha Klimaszewski & Webster in Klimaszewski et al., 2018a: 2

Type species: Atheta meduxnekeagensis Webster & Klimaszewski in Klimaszewski et al., 2018 Hypsostiba Bernhauer, 1929a: 200 [*]

Type species: Atheta dampfi Bernhauer, 1929

Ischnoderoecia Scheerpeltz, 1974: 249

Type species: Atheta lunaris Scheerpeltz, 1974

Kamptodota Pace, 2009: 274

Type species: Atheta papuaimitatrix Pace, 2009

Kladobrachydotina Pace, 2008a: 302 [*]

Type species: Atheta inflatheca Pace, 2008

Lamiota Casey, 1910a: 17

Type species: Atheta keeni Casey, 1910

Lampratheta Scheerpeltz, 1965b: 334

Type species: Atheta malaisei Scheerpeltz, 1965

Libanostiba Scheerpeltz, 1929: 227

Type species: Atheta ebneri Scheerpeltz, 1929

Lobodota Pace, 2009: 274

Type species: Atheta nabirensis Pace, 2009

Metadimetrota Klimaszewski & Winchester, 2002: 31

Type species: Atheta cheersae Klimaszewski, 2002

Microdota Mulsant & Rey, 1873d: 160 [*]

Type species: Microdota sericea Mulsant & Rey, 1873

Anatelloleptusa Scheerpeltz, 1976c: 50 [ Smetana 2004: 384]

Type species: Leptusa himalayica Scheerpeltz, 1976

Dicolyota Pace, 2004: 359 [nomen nudum, synonymized by Schülke & Smetana 2015: 542]

Type species: Atheta praelata Pace, 1986

Dicolyota Pace, 1998: 924 [synonymized by Pace 2004: 359]

Type species: [no type species designated]

Heteronoma Mulsant & Rey, 1873b: 4 [synonymized by Bernhauer & Scheerpeltz 1926: 630]

Type species: Homalota luctuosa Mulsant & Rey, 1853

Heterophaena Lynch-Arribálzaga, 1884: 45 [synonymized by Bernhauer & Scheerpeltz 1 926: 630]

Type species: Heterophaena palliditarsis Lynch-Arribálzaga, 1884

Metamischa Peyerimhoff, 1938: 65 [synonymized by Pace 1990c: 915]

Type species: Homalota scrobicollis Kraatz, 1859

Meoticaemorpha Scheerpeltz, 1976c: 66 [ Smetana 2004: 384]

Type species: Meoticaemorpha janetscheki Scheerpeltz, 1976

Physadota Pace, 1990c: 907 [synonymized by Pace 2004c: 359]

Type species: Atheta subscabra Cameron, 1939

Sonomota Casey, 1911: 158 [synonymized by Lohse & Smetana 1988: 270]

Type species: Sipalia lippa Casey, 1911

Microdotina Scheerpeltz, 1974: 254

Type species: Atheta xanthopyga Scheerpeltz, 1974

Nemota Casey, 1910a: 56

Type species: Atheta paganella Casey, 1910

Niphetodroma Scheerpeltz, 1947: 288

Type species: Niphetodroma obsolescens Scheerpeltz, 1947

Oligatheta Bernhauer & Scheerpeltz, 1926: 605 [*]

Type species: Atheta cordillerana Bernhauer, 1921

Micratheta Bernhauer, 1921b: 179 [synonymized by Bernhauer & Scheerpeltz 1926: 605]

Type species: Atheta cordillerana Bernhauer, 1921

Oreostiba Ganglbauer, 1895: 219

Type species: Homalota tibialis Heer, 1839

Agaphygra Tottenham, 1949c: 78 [ Smetana 2004: 31]

Type species: Homalota subglabra Sharp, 1869

Orientatheta Sawada, 1985: 99

Type species: Atheta chinkiangensis Bernhauer, 1939

Oxypodera Bernhauer, 1915d: 185

Type species: Atheta kilimandjarensis Bernhauer, 1915

Pachyatheta Münster, 1925: 11

Type species: Homalota cribrate Kraatz, 1856

Paradota Sawada, 1989: 274

Type species: Atheta liberta Sawada, 1989

Parameotica Ganglbauer, 1895: 228 [*]

Type species: Homalota laticeps Thomson, 1856

Parametaxya Jeannel & Paulian, 1945: 106

Type species: Atheta jeanneliana Bernhauer, 1944

Paramidobia Bernhauer, 1908: 356 [*]

Type species: Amidobia longiceps Bernhauer, 1908

Parataxicera Brundin, 1943: 27

Type species: Atheta moczarskii Bernhauer, 1915

Peliolurga Tottenham, 1939b: 228

Type species: Bolitochara luridipennis Mannerheim, 1830

Pelurga Mulsant & Rey, 1873b: 577 [synonymized by Tottenham 1939b: 228]

Type species: Bolitochara luridipennis Mannerheim, 1830

Pelmatheta Scheerpeltz, 1965b: 335

Type species: Atheta crassitarsis Scheerpeltz, 1965

Philacrotona Pace, 1991c: 140

Type species: Atheta newar Pace, 1991

Polydota Pace, 1983d: 310 [*]

Type species: Atheta uhligi Pace, 1983

Poromicrodota Pace, 1990c: 907

Type species: Atheta subamicula Cameron, 1939

Pseudhygroecia Jeannel & Paulian, 1945: 104

Type species: Atheta elgonensis Bernhauer, 1944

Pseudobessobia Bernhauer, 1921b: 177 [*]

Type species: Atheta weiseri Bernhauer, 1921

Pseudohygroecia Bernhauer, 1929c: 189

Type species: Atheta crisiana Bernhauer, 1929

Pseudoleptonia Bernhauer, 1934c: 507 [*]

Type species: Atheta polyporina Bernhauer, 1934

Pseudophilygra Bernhauer, 1929c: 190

Type species: Atheta holtzi Bernhauer, 1929

Pseudota Casey, 1910a: 114

Type species: Pseudota dissensa Casey, 1910

Pseudothinoecia Bernhauer, 1899: 20

Type species: Atheta puellaris Bernhauer, 1899

Rhagocneme Münster, 1923: 206

Type species: Atheta fractipes Münster, 1923

Sepedomicra Sawada, 1985: 97

Type species: Atheta mycetophaga Cameron, 1920

Sipalatheta Pace, 1993: 92

Type species: Atheta difformis Pace, 1993

Spelaeolla Rambousek, 1915b: 31

Type species: Atheta absoloni Rambousek, 1915

Stictatheta Cameron, 1939a: 5

Type species: Atheta quadripunctula Cameron, 1939

Stilbatheta Scheerpeltz, 1962a: 596

Type species: Atheta speculiventris Scheerpeltz, 1962

Strobilocera Ganglbauer, 1895: 191

Type species: Homalota capitulata Eppelsheim, 1880

Suensonia Bernhauer, 1936d: 314

Type species: Atheta obsoletepunctata Bernhauer, 1936

Synaloconota Scheerpeltz, 1974: 247

Type species: Atheta cascadensis Scheerpeltz, 1974

Tachynota Bernhauer, 1901b: 113

Type species: Atheta thinodromoides Bernhauer, 1901

Tetropla Mulsant & Rey, 1873b: 492

Type species: Aleochara nigritula Gravenhorst, 1802

Thinobaena Thomson, 1859: 39

Type species: Aleochara quisquiliarum Gyllenhal, 1810

Parabessobia Scheerpeltz, 1958: 32 [synonymized by G. Benick 1973: 187]

Type species: Atheta serricauda Eppelsheim, 1878

Traumoecia Mulsant & Rey, 1873b: 631

Type species: Aleochara excavata Gyllenhal, 1827

Trochanterella Brundin, 1953: 428

Type species: Atheta cribripennis J. Sahlberg, 1890

Tropatheta Bernhauer, 1927a: 81

Type species: Atheta gestroi Bernhauer, 1927

Umbala Blackwelder, 1952: 402

Type species: Atheta mimetica Cameron, 1939

Stictatheta Cameron, 1939b: 336 [synonymized by Blackwelder 1952: 402]

Type species: Atheta mimetica Cameron, 1939

Xenota Mulsant & Rey, 1873b: 397 [*]

Type species: Homalota myrmecobia Kraatz, 1856

Mycota Mulsant & Rey, 1873b: 502 [ Newton & Thayer 2003]

Type species: Homalota humeralis Kraatz, 1856

Xestota Bernhauer, 1908: 361 [*]

Type species: Atheta biarmata Bernhauer, 1908

353 species are included in this genus from Latin America.

Characters/Description: Lynch-Arribálzaga 1884: 50 (characters of subgenus Microdota ); Fenyes 1920: 181 (characters), 201 (characters as Synaptina ), 210 (characters subgenus Atheta ), 238 (characters as Iotota ), 247 (characters as Rhodeota ); Lohse et al. 1990: 125 (characters in key), 126 (characters in key of Bessobia ), 126 (characters in key of Dimetrota ), 178 (characters of Dimetrota ), 188 (characters); Ashe 2002b: 128 (characters in key); Pace 2008a: 281 (characters in key); Aballay et al. 2014: 103 (characters in key).

Key: Fenyes 1920: 182 (subgenera); Pace 1988: 88 (some species of Chile); Pace 1996c: 652 (some species of Ecuador); Klimaszewski & Peck, 1998: 236 (species of the Galapagos Isl.); Pace 2000a: 428 (some species of Chile); Pace 2008a: 294 (subgenera of Ecuador and Peru), 302 (species of the subgenus Acrotona of Ecuador and Peru), 305 (species of the subgenus Xestota of Ecuador and Peru), 307 (species of the subgenus Microdota of Ecuador and Peru), 310 (species of the subgenus Kladobrachydotina of Ecuador and Peru), 312 (species of the subgenus Datomicra of Ecuador and Peru), 317 (species of the subgenus Dimetrota of Ecuador and Peru), 319–321 (species of the subgenus Pseudobessobia of Ecuador and Peru); Newton & Peck 2006: 139 (species of the Galapagos Isl.).

Distribution.

West Indies: Cuba, Cayman Is., Jamaica, Haiti, Puerto Rico, US Virgin Isl., British Virgin Isl., Guadeloupe, St. Lucia, St. Vincent & Grenadines, Grenada.

Central America: Mexico, Guatemala, Belize, Nicaragua, Costa Rica, Panama.

South America: Colombia, Venezuela, Suriname, French Guiana, Ecuador, Galapagos Isl., Peru, Bolivia, Brazil, Paraguay, Chile, Argentina.

Other regions: Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian, Oceanic.

0305. Brundinia Tottenham, 1949c: 78

Type species: Metaxya apricans Mulsant & Rey, 1873

Metaxya Mulsant & Rey, 1873d: 181 [synonymized by Tottenham 1949c: 78]

Type species: Metaxya apricans Mulsant & Rey, 1873

4 species are included in this genus from South America.

Characters/Description: Fenyes 1920: 197 (characters as Metaxya ).

Distribution.

South America: Peru, Argentina.

Other regions: Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian.

0306. Dalotia Casey, 1910a: 106

Type species: Dimetrota pectorina Casey, 1910

1 species is included in this genus from the West Indies and South America.

Key: Newton & Peck 2006: 139 (species of the Galapagos Isl.).

Distribution.

West Indies: Dominican Republic.

South America: Ecuador, Galapagos Isl., Bolivia, Brazil, Chile, Argentina.

Other regions: Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian.

0307. Dochmonota Thomson, 1859: 40

Type species: Homalota funebris Thomson, 1856

1 species is included in this genus from South America.

Characters/Description: Fenyes 1920: 189 (characters).

Distribution.

South America: Argentina.

Other regions: Nearctic, Palaearctic.

0308. Homia Blackwelder, 1952: 190

Type species: Daya occipitalis Fauvel, 1878

Daya Fauvel, 1878d: 147 [synonymized by Blackwelder 1952: 118]

Type species: Daya occipitalis Fauvel, 1878

1 species is included in this genus from South America.

Characters/Description: Fenyes 1920: 230 (characters as Daya ).

Distribution.

South America: Brazil.

Other regions: Nearctic, Palaearctic.

0309. Hydrosmecta Thomson, 1858: 33

Type species: Homalota longula Heer, 1839

Actocharina Bernhauer in Ganglbauer, 1907: 185 [synonymized by Elven et al. 2012: 633]

Type species: Atheta leptotyphloides Bernhauer in Ganglbauer, 1907

Doliponta Blackwelder, 1952: 132 [synonymized by Blackwelder 1952: 132]

Type species: Lipodonta veris Fenyes, 1921

Hydrosmectina Ganglbauer, 1895: 232 [ Smetana 2004: 407]

Type species: Homalota subtilissima Kraatz, 1854

Lipodonta Fenyes, 1921a: 24 [synonymized by Blackwelder 1952: 132]

Type species: Lipodonta veris Fenyes, 1921

Thinoecia Mulsant & Rey, 1873d: 184 [synonymized by Blackwelder 1952: 386]

Type species: Thinoecia libitina Mulsant & Rey, 1873

Thinaecia Mulsant & Rey, 1873b: 5 [synonymized by Blackwelder 1952: 386]

Thinaecia Mulsant & Rey, 1874c: 37 [synonymized by Blackwelder 1952: 386]

Thinoecia Mulsant & Rey, 1874a: 38 [synonymized by Blackwelder 1952: 386]

Thinoecia Mulsant & Rey, 1875a: 260 [synonymized by Blackwelder 1952: 386]

Thinoecia Mulsant & Rey, 1875c: 234 [synonymized by Bernhauer & Scheerpeltz 1 926: 605]

7 species are included in this genus from Central and South America. Characters/Description: Fenyes 1920: 174 (characters as Actocharina ), 236 (characters), 237 (characters as Hydrosmectina ); Lohse et al. 1990: 125 (characters in key), 147 (characters); Pace 2008a: 280 (characters in key). Distribution. Central America: Mexico, Guatemala, Panama. South America: Ecuador, Chile, Argentina. Other regions: Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian, Oceanic.

0310. Nehemitropia Lohse, 1971: 81

Type species: Staphylinus sordidus Marsham, 1802

1 species is included in this genus from South America.

Distribution.

South America: Peru, Bolivia, Brazil, Paraguay, Uruguay, Chile, Argentina.

Other regions: Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian.

0311. Notothecta Thomson, 1858: 33

Type species: Aleochara flavipes Gravenhorst, 1806

Subgenera:

Kraatzia Saulcy, 1862: 289

Type species: Kraatzia attophila Saulcy, 1862

Notothecta Thomson, 1858: 33

Type species: Aleochara flavipes Gravenhorst, 1806

Notothectina Bernhauer, 1912d: 77 [*]

Type species: Notothecta attae Bernhauer, 1912

2 species are included in this genus from South America.

Characters/Description: Fenyes 1920: 240 (characters), 241 (characters of subgenera Kraatzia and Notothecta ), 242 (characters of subgenus Notothectina ).

Key: Fenyes 1920: 241 (subgenera).

Distribution.

South America: Argentina.

Other regions: Palaearctic, Oriental.

0312. Philhygra Mulsant & Rey, 1873d: 160

Type species: Microdota perdubia Mulsant & Rey, 1873

Amphibitherion Notman, 1921: 155 [ Seevers 1978: 123]

Type species: Amphibitherion demissum Notman, 1921

Homalotusa Casey, 1906: 340 [ Smetana 2004: 390]

Type species: Homalotusa helenica Casey, 1906

Hygroecia Mulsant & Rey, 1873d: 187 [ Schülke & Smetana 2015: 547]

Type species: Hygroecia parca Mulsant & Rey, 1873

Phryogora Mulsant & Rey, 1873b: 625 [ Schülke & Smetana 2015: 548]

Type species: Homalota hygrobia Thomson, 1856

Plagiarthrina Keys, 1920: 131 [ Smetana 2004: 390]

Type species: Metaxya fordhamiana Keys, 1920

Valenusa Casey, 1906: 342 [ Newton & Thayer 2003]

Type species: Valenusa parallela Casey, 1906

2 species are included in this genus from South America.

Characters/Description: Fenyes 1920: 200 (characters); Lohse et al. 1990: 126 (characters in key), 167 (characters). Distribution.

South America: Colombia, Brazil.

Other regions: Neartic, Palaearctic, Ethiopian.

0313. Platyola Mulsant & Rey, 1875e: 249

Type species: Homalota fusicornis Mulsant & Rey, 1853

Australoplatyola Scheerpeltz, 1959b: 10 [nomen nudum]

Type species: Platyola antarctica Bernhauer, 1920

Neoplatyola Pace, 1996b: 399 [nomen nudum]

Platyola Mulsant & Rey, 1875d: 411 [synonymized by Blackwelder 1952: 313]

3 species are included in this genus from South America.

Characters/Description: Fenyes 1920: 376 (characters).

Distribution.

South America: Colombia, Peru, Bolivia, Brazil.

Other regions: Palaearctic, Oriental.

0314. Pycnota Mulsant & Rey, 1873b: 377

Type species: Homalota paradoxa Mulsant & Rey, 1861

1 species is included in this genus from South America.

Characters/Description: Fenyes 1920: 246 (characters).

Key: Pace, 1988: 86 (some species of Chile).

Distribution.

South America: Brazil.

Other regions: Palaearctic.

0315. Schistoglossa Kraatz, 1856b: 344 Type species: Homalota viduata Erichson, 1837 Hölzelia Scheerpeltz, 1952b: 765 [ Smetana 2004: 416] Type species: Hoelzelia caricis Scheerpeltz, 1952 Protoskiusa Bernhauer, 1900b: 200 [synonymized by Bernhauer & Scheerpeltz 1926: 591] Type species: Protoskiusa paradoxa Bernhauer, 1900

1 species is included in this genus from South America.

Characters/Description: Fenyes 1920: 248 (characters).

Distribution.

South America: Ecuador, Peru.

Other regions: Nearctic, Palaearctic, Oriental.

0316. Stethusa Casey, 1910a: 4

Type species: Atheta klimschi Bernhauer, 1909

3 species are included in this genus from the West Indies and South America.

Characters/Description: Gusarov 2003c: 5 (characters).

Key: Gusarov 2003c: 12 (species of the genus); Newton & Peck 2006: 139 (species of the Galapagos Isl.). Distribution.

West Indies: US Virgin Isl., Guadeloupe, St. Lucia., Grenada.

South America: Venezuela, French Guiana, Galapagos Isl., Bolivia, Brazil, Paraguay, Uruguay, Argentina.

Other regions: Nearctic, Palaearctic?.

Figures

FIGURES 40–51. Habitus; Habrocerinae (40), Aleocharinae (41–51); 40, after Solier 1849; 41, Brazil: state of Paraná (MUSM); 42, Chile: region of Antofagasta (MUSM); 43, after Pace 2015e; 44, after Ashe 2002a; 45, after Pace 2014a; 46, after Pace 2012; 47, after Y.H. Kim & Ahn 2015; 48, after Pace 2014c; 49, Chile: region of Araucanía (MUSM); 50, after Irmler 2011; 51, after Caron & Ribeiro-Costa 2008; slightly modified. Scale bars: 1 mm.