Brachyglutina Raffray, 1904b

Asenjo, Angélico, Irmler, Ulrich, Klimaszewski, Jan, Chandler, Donald S., Fierros-López, Hugo Eduardo & Vieira, Juliana S., 2019, Staphylinidae (Insecta: Coleoptera) in Latin America: synopsis, annotated catalog, diversity and distribution, Zootaxa 4621 (1), pp. 1-406 : 57-65

Subtribe Brachyglutina Raffray, 1904b

0174. Achilia Reitter, 1890: 212

Type species: Bryaxis valdiviensis Reitter, 1883

Byraxis Raffray, 1890b: 123 [synonymized by Reitter 1890: 212]

Type species: Bryaxis valdiviensis Reitter, 1883

Clermontodes Jeannel, 1950: 317 [synonymized by Besuchet 1986: 259]

Type species: Clermontodes insularis Jeannel, 1950

68 species are known from and are restricted to South America. Achilia insularis (Jeannel) described from France was mislabeled.

Characters/Description: Raffray 1904a: 110 (characters in key as Byraxis ), 113 (characters in key); Raffray 1908a: 193 (characters in key as Byraxis ), 195 (characters in key), 220 (characters); O. Park 1942: 126 (characters in key), 154 (characters); O. Park 1944: 241 (characters in key); Jeannel 1962b: 391 (characters in key), 396 (characters); H. Franz 1996: 114 (characters); Kurbatov & Sabella 2015: 304 (characters); Sabella et al. 2017: 120 (charactes of Achilia crassicornis species group), 123 (characters of Achilia tumidifrons group), 127 (characters of Achilia bifossifrons group), 132 (characters of Achilia lobifera group); Kurbatov et al. 2018: 166 (characters of Achilia frontalis species group).

Key: Reitter 1885b: 324 (species of the genus as Bryaxis ); Raffray 1904a: 134 (species groups); O. Park 1942: 155 (species groups); Jeannel 1962b: 397 (species groups), 399, 402, 405, 407, 409, 421, 424, 425, 431, 435, 438 (species in the groups).

Distribution.

South America: Chile, Argentina.

0175. Achillidia Jeannel, 1962b: 391

Type species: Bryaxis bituberculata Reitter, 1885

Only one species is known from and is restricted to South America.

Characters/Description: Jeannel 1962b: 390 (characters in key).

Distribution.

South America: Chile.

0176. Achilliotes Jeannel, 1962b: 445

Type species: Achilia brevicornis Raffray, 1904

Only one species is known from and is restricted to South America.

Characters/Description: Jeannel 1962b: 391 (characters in key).

Distribution.

South America: Chile.

0177. Anchylarthron Brendel, 1887: 208

Type species: Decarthron cornutum Brendel, 1865

Verticinotus Brendel, 1888: 300 [synonymized by Newton & Chandler 1989: 45]

Type species: Decarthron cornutum Brendel, 1865

1 species is included in this genus from Central America. Characters/Description: Raffray 1904a: 111 (characters in key); Raffray 1908a: 194 (characters in key), 212

(characters); Becker & Sanderson 1953a: 403 (characters in key); Chandler 2002: 53 (characters in key). Distribution. Central America: Mexico. Other regions: Nearctic.

0178. Atacamia H. Franz, 1996: 111

Type species: Atacamia paludosa H. Franz, 1996

Only one species is known from and is restricted to South America.

Distribution.

South America: Chile.

0179. Berdura Reitter, 1882b: 206

Type species: Berdura excisula Reitter, 1883

3 species are known from and are restricted to the West Indies and Central America.

Characters/Description: Raffray 1904a: 110 (characters in key); Raffray 1908a: 193 (characters in key), 205 (characters); O. Park 1942: 124 (characters in key), 163 (characters).

Key: O. Park 1942: 164 (species of the genus); O. Park et al. 1976: 49 (species of the West Indies).

Distribution.

West Indies: Cuba, US Virgin Isl.

Central America: Panama.

0180. Brachygluta Thomson, 1859: 54

Type species: Pselaphus fossulatus Reichenbach, 1816

Subgenera:

Brachygluta Thomson, 1859: 54 [*]

Type species: Pselaphus fossulatus Reichenbach, 1816

Brachyglutina Jeannel, 1950: 286 [synonymized by Karaman 1961: 128]

Type species: Bryaxis globulicollis Mulsant & Rey, 1861

Brachyglutodes Jeannel, 1956: 97 [synonymized by Karaman 1961: 128]

Type species: Bryaxis leprieuri Saulcy, 1876

Dicrobiella Jeannel, 1950: 315 [synonymized by Jeannel 1956: 82]

Type species: Bryaxis dentiventris Saulcy, 1876

Dicrobionoma Jeannel, 1956: 97 [ Newton & Chandler 1989: 42]

Type species: Bryaxis caligata Saulcy, 1876

Neobrachygluta Kaszab & Székessy, 1953: 475 [synonymized by Besuchet 1963: 27]

Type species: Brachygluta dudichi Kaszab & Székessy, 1953

Nisa Casey, 1886b: 182

Type species: Bryaxis cavicornis Brendel, 1865

5 species are included in this genus from Central America.

Key: Raffray 1904a: 141 (species group); O. Park 1942: 158 (species of the genus).

Distribution. South America: Brazil.

0184. Bryaxinella Jeannel, 1962b: 394

Type species: Bryaxinella nodicornis Jeannel, 1962

Only one species is known from and is restricted to South America.

Characters/Description: Jeannel 1962b: 391 (characters in key).

Distribution.

South America: Argentina.

0185. Bunoderus Raffray, 1904a: 151

Type species: Bunoderus carinicollis Raffray, 1904

18 species are known from and are restricted to Central and South America.

Characters/Description: Raffray 1904a: 115 (characters in key); Raffray 1908a: 197 (characters in key), 231 (characters); O. Park 1942: 125 (characters in key), 154 (characters); O. Park 1944: 248 (characters in key); Chandler 1983a: 204 (characters); Chandler 2002: 54 (characters in key).

Key: O. Park 1942: 154 (species of the genus); O. Park 1944: 251 (Mexican species); Chandler 1983a: 205 (species of the genus, males).

Distribution.

Central America: Mexico, Nicaragua, Panama.

South America: Brazil.

0186. Byraxorites Jeannel, 1962b: 389

Type species: Bryaxorites alticola Jeannel, 1962

Only one species is known from and is restricted to South America.

Characters/Description: Jeannel 1962b: 387 (characters in key).

Distribution.

South America: Chile.

0187. Bythinogaster L.W. Schaufuss, 1887: 111

Type species: Bythinogaster simplex L.W. Schaufuss, 1887

5 species are known from and are restricted to the West Indies.

Characters/Description: Raffray 1904a: 109 (characters in key); Raffray 1908a: 193 (characters in key), 203 (characters); O. Park 1942: 124 (characters in key), 203 (characters); O. Park et al. 1976: 53 (characters).

Key: O. Park et al. 1976: 53 (species of the West Indies).

Distribution.

West Indies: Cuba, Jamaica, "Hispaniola".

0188. Cryptorhinula L.W. Schaufuss, 1887: 150 , 163

Type species: Cryphorhinula nodifera L.W. Schaufuss, 1887

5 species are known from and are restricted to South America.

Characters/Description: Raffray 1890b: 124 (characters); Raffray 1897: 257 (characters in key); Raffray 1904a: 112 (characters in key); Raffray 1908a: 194 (characters in key), 215 (characters); O. Park 1942: 125 (characters in key), 179 (characters).

Key: O. Park 1942: 180 (species of the genus).

Distribution.

South America: Venezuela, Brazil.

0189. Drasinus Raffray, 1904a: 148

Type species: Drasinus binodulus Raffray, 1904

Subgenera:

Drasinus Raffray, 1904a: 148 [*]

Type species: Drasinus binodulus Raffray, 1904

Paradrasinus Newton & Chandler, 1989: 43 [*]

Type species: Drasinus hirsutus Fletcher, 1928

Paradrasinus O. Park, 1942: 182 [nomen nudum, validated by Newton & Chandler 1989: 43]

Type species: [no type species designated]

4 species are known from and are restricted to Central and South America.

Characters/Description: Raffray 1904a: 114 (characters in key); Raffray 1908a: 196 (characters in key), 226 (characters); O. Park 1942: 124 (characters in key), 181 (characters); O. Park 1944: 242 (characters in key); Becker & Sanderson 1953a: 403 (characters in key); Becker & Sanderson 1953b: 437 (characters); Chandler 2002: 53 (characters in key).

Key: O. Park 1942: 181 (species of the genus); O. Park 1944: 251 (Mexican species).

Distribution.

Central America: Mexico, Belize, Honduras, Nicaragua, Costa Rica, Panama.

South America: Suriname, Brazil.

0190. Ectopocerus Raffray, 1904a: 149

Type species: Decarthron verticicornis Reitter, 1885

Only one species is known from and is restricted to South America.

Characters/Description: Raffray 1904a: 114 (characters in key); Raffray 1908a: 196 (characters in key), 226 (characters); O. Park 1942: 124 (characters in key), 185 (characters); Jeannel 1962b: 391 (characters in key), 443 (characters).

Distribution.

South America: Chile.

0191. Eremomus Raffray, 1904a: 177

Type species: Eremomus obesus Raffray, 1904

2 species are known from and are restricted to South America.

Characters/Description: Raffray 1904a: 115 (characters in key); Raffray 1908a: 197 (characters in key), 242 (characters); O. Park 1942: 124 (characters in key), 133 (characters).

Key: O. Park 1942: 134 (species of the genus).

Distribution.

South America: Bolivia, Brazil.

0192. Globa Raffray, 1887: 37 ( Fig. 23 )

Type species: Globa longipes Raffray, 1887

3 species are known from and are restricted to Central and South America. Characters/Description: Raffray 1897: 256 (characters in key), 258 (characters); Raffray 1904a: 110 (characters in key); Raffray 1908a: 193 (characters in key), 204 (characters); O. Park 1942: 124 (characters in key), 202 (characters); Chandler 2002: 53 (characters in key). Key: O. Park 1942: 203 (species of the genus). Distribution. Central America: Panama. South America: Colombia, Venezuela, Bolivia.

0193. Leptachillia Jeannel, 1962b: 446

Type species: Leptachillia laevissima Jeannel, 1962

2 species are known from and are restricted to South America.

Characters/Description: Jeannel 1962b: 391 (characters in key).

Distribution.

South America: Chile.

0194. Mallecoa H. Franz, 1996: 110

Type species: Mallecoa abnormis H. Franz, 1996

Only one species is known from and is restricted to South America.

Distribution.

South America: Chile.

0195. Mitona Raffray, 1904a: 129

Type species: Mitona quadraticeps Raffray, 1904

4 species are known from and are restricted to South America.

Characters/Description: Raffray 1904a: 111 (characters in key); Raffray 1908a: 194 (characters in key), 213 (characters); O. Park 1942: 125 (characters in key), 175 (characters); O. Park 1945b: 394 (characters).

Key: O. Park 1942: 176 (species of the genus); O. Park 1945b: 395 (species of the genus).

Distribution.

South America: Colombia, Venezuela, Bolivia.

0196. Nisaxis Casey, 1886b: 183

Type species: Bryaxis tomentosa Aubé, 1833

Dranisaxa O. Park, 1945b: 391 [synonymized by Chandler 2000: 174]

Type species: Dranisaxa gloydi O. Park, 1945

3 species are included in this genus from the West Indies and Central America.

Characters/Description: Raffray 1904a: 112 (characters in key); Raffray 1908a: 195 (characters in key), 219 (characters); Chandler 2002: 53 (characters in key).

Distribution.

West Indies: Cayman Is.

Central America: Mexico.

Other regions: Nearctic.

0197. Nondulia Newton & Chandler, 1989: 43

Type species: Bryaxis convexa L.W. Schaufuss, 1880

Nodulina Raffray, 1904a: 115 [synonymized by Newton & Chandler 1989: 43]

Type species: Bryaxis convexa L.W. Schaufuss, 1880

Only one species is known from and is restricted to South America.

Characters/Description: Raffray 1904a: 115 (characters in key as Nondulia ); Raffray 1908a: 197 (characters in key as Nondulia ), 232 (characters as Nondulia ); O. Park 1942: 125 (characters in key), 153 (characters).

Distribution.

South America: Guyana, Suriname, Bolivia, Brazil.

0198. Panabachia O. Park, 1942: 153

Type species: Bryaxis vulnerata Sharp, 1887

2 species are known from and are restricted to Central America.

Characters/Description: O. Park 1942: 125 (characters in key); O. Park 1945b: 389 (characters); Chandler 2002: 53 (characters in key).

Distribution.

Central America: Mexico, Guatemala, Panama.

0199. Parachillia H. Franz, 1996: 112

Type species: Parachillia longicornis H. Franz, 1996

3 species are known from and are restricted to South America.

Distribution.

South America: Chile.

0200. Pseudachillia Jeannel, 1963: 367

Type species: Pseudachillia bicolor Jeannel, 1963

2 species are known from and are restricted to South America.

Distribution. South America: Chile.

0201. Pseudachillidia H. Franz, 1996: 112

Type species: Pseudachillidia andina H. Franz, 1996

Only one species is known from and is restricted to South America.

Distribution.

South America: Chile.

0202. Raxybis Raffray, 1908a: 222

Type species: Raxybis nodosa Raffray, 1908

2 species are known from and are restricted to South America.

Characters/Description: Raffray 1908a: 195 (characters in key); O. Park 1942: 126 (characters in key), 157 (characters); Jeannel 1962b: 388 (characters in key), 389 (characters).

Key: O. Park 1942: 157 (species of the genus).

Distribution.

South America: Argentina.

0203. Reichenbachia Leach, 1826: 451 Type species: Bryaxis juncorum Leach, 1817 Dierobia Thomson, 1859: 54 [synonymized by Reitter 1885c: 68] Type species: Bryaxis juncorum Leach, 1817 Reichenbachius Casey, 1906: 359 [synonymized by Newton & Chandler 1989: 44] Type species: Bryaxis juncorum Leach, 1817

Key: Raffray 1904a: 125 (species group); O. Park 1943: 191 (some species of the genus); O. Park 1942: 167–170 (species group and species); O. Park 1944: 251 (Mexican species); Becker & Sanderson 1953a: 427 (species of Honduras); O. Park et al. 1976: 51 (species of the West Indies).

Distribution.

West Indies: Jamaica, Guadeloupe, St. Vincent & Grenadines, Grenada.

Central America: Mexico, Guatemala, Belize, Honduras, Panama.

South America: Bolivia, Brazil, Paraguay.

Other regions: Nearctic.

0206. Silillicus Lucas, 1920: 504

Type species: Phoberus armatus Raffray, 1904

Phoberus Raffray, 1904a: 176 [synonymized by Lucas 1920: 504]

Type species: Phoberus armatus Raffray, 1904

Only one species is known from and is restricted to South America.

Characters/Description: Raffray 1904a: 115 (characters in key as Phoberus ), 176 (characters as Phoberus ); Raffray 1908a: 197 (characters in key as Phoberus ), 241 (characters as Phoberus ); O. Park 1942: 124 (characters in key as Phoberus ), 134 (characters as Phoberus ).

Distribution.

South America: Bolivia.

0207. Strombopsis Raffray, 1904a: 133

Type species: Strombopsis breviventris Raffray, 1904

Only one species is known from and is restricted to South America.

Characters/Description: Raffray 1904a: 112 (characters in key); Raffray 1908a: 195 (characters in key), 218 (characters); O. Park 1942: 126 (characters in key); O. Park 1944: 241 (characters in key).

Distribution.

South America: Brazil.

0208. Xybarida Raffray, 1897: 260

Type species: Xybarida clavata Raffray, 1897

5 species are known from and are restricted to Central and South America. Characters/Description: Raffray 1897: 257 (characters in key); Raffray 1904a: 112 (characters in key); Raffray 1908a: 195 (characters in key), 217 (characters); O. Park 1942: 126 (characters in key), 160 (characters); O. Park 1944: 249 (characters in key); Chandler 2002: 53 (characters in key). Key: O. Park 1942: 160 (species of the genus); O. Park 1944: 251 (Mexican species). Distribution. Central America: Mexico, Guatemala, Panama. South America: Brazil.

0209. Xybaris Reitter, 1882a: 140

Type species: Xybaris spiniceps Reitter, 1882

7 species are known from and are restricted to South America.

Characters/Description: Reitter 1882a: 139 (characters in key); Raffray 1897: 257 (characters in key); Raffray 1904a: 111 (characters in key); Raffray 1908a: 194 (characters in key), 214 (characters); O. Park 1942: 125 (characters in key), 176 (characters).

Distribution.

South America: Colombia, Brazil.

Figures

FIGURES 16–27. Habitus; Micropeplinae (16–18), Neophoninae (19), Pselaphinae (20–27); 16, after Campbell 1968; 17, after Campbell 1973c; 18, after Tronquet & Degallier 2010; 19 Chile: region of Los Lagos (VMDC); 20, after Reitter 1883b; 21, after Reitter 1889; 22, after Bryant 1915; 23, after Raffray 1887; 24, after Asenjo 2016; 25, after Asenjo et al. 2017; 26, after O. Park 1952b; 27, after Bruch 1929; slightly modified. Scale bars: 1 mm.