Bolitocharina Thomson, 1859

Asenjo, Angélico, Irmler, Ulrich, Klimaszewski, Jan, Chandler, Donald S., Fierros-López, Hugo Eduardo & Vieira, Juliana S., 2019, Staphylinidae (Insecta: Coleoptera) in Latin America: synopsis, annotated catalog, diversity and distribution, Zootaxa 4621 (1), pp. 1-406 : 134-140

publication ID	https://doi.org/10.11646/zootaxa.4621.1.1
publication LSID	lsid:zoobank.org:pub:3D77F329-9BB8-4231-90E4-6A9E0250FCDF
persistent identifier	https://treatment.plazi.org/id/03978788-B9F1-FF4E-C1E3-6ABDAC509B6B
treatment provided by	Plazi (2019-06-26 09:05:59, last updated 2024-11-25 23:02:34)
scientific name	Bolitocharina Thomson, 1859
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Treatment

Subtribe Bolitocharina Thomson, 1859 View in CoL

0445. Bolitochara Mannerheim, 1830: 75

Type species: Aleochara pulchra Gravenhorst, 1806

Agaribiota Bierig, 1937b: 279 [synonymized by Blackwelder 1952: 41]

Type species: Bolitochara cinctigastra Bierig, 1937

Ditropalia Casey, 1906: 263 View in CoL [synonymized by Seevers 1978: 164]

Type species: Bolitochara bella Märkel, 1844

3 species are included in this genus from the West Indies and South America.

Characters/Description: Fenyes 1920: 114 (characters as Ditropalia ); Coiffait & Sáiz 1967: 65 (characters in key), 79 (characters); Ashe 1992: 365 (characters in key), 367 (characters).

Key: Coiffait & Sáiz 1967: 80 (Chilean species).

Distribution.

West Indies: Cuba.

South America: Colombia, Argentina.

Other regions: Palaearctic, Oriental, Australian, Oceanic.

0446. Habramicrada Pace, 2007: 225 ( Fig. 58 View FIGURES 52–63 )

Type species: Habramicrada boliviensis Pace, 2007

4 species are known from and are restricted to South America.

Distribution.

South America: Peru, Bolivia, Brazil.

0447. Leptusa Kraatz, 1856b: 60

Type species: Bolitochara pulchella Mannerheim, 1830

Subgenera:

Adexiopisalia Pace, 1982c: 585

Type species: Leptusa hummleri Bernhauer, 1910

Adoxopisalia Pace, 1989: 130

Type species: Leptusa opaca Casey, 1893

Afroleptusa Pace, 1983c: 62

Type species: Leptusa sparsa Fauvel, 1898

Akratopisalia Pace, 1996a: 29

Type species: Leptusa cribrata Pace, 1996

Amalopisalia Pace, 1983c: 97

Type species: Leptusa janetscheki Scheerpeltz, 1956

Anisopisalia Pace, 1989: 214

Type species: Leptusa luigionii Bernhauer, 1935

Anomopisalia Pace, 1983c: 82

Type species: Leptusa granulicauda Eppelsheim, 1890

Anosiopisalia Pace, 1995: 247

Type species: Leptusa nemoricultrix Pace, 1995

Aphaireleptusa Pace, 1996a: 30

Type species: Leptusa anmashanensis Pace, 1996

Aleteleptusa Pace, 1997a: 753 [ Smetana 2004: 429]

Type species: Leptusa chinensis Pace, 1997

Mimumenepisalia Pace, 1997a: 752 [ Smetana 2004: 429]

Type species: Leptusa gonggamontis Pace, 1997

Baldopisalia Pace, 1983e: 130

Type species: Leptusa veronensis Pace, 1983

Boreoleptusa Pace, 1989: 82

Type species: Leptusa norvegica Strand, 1941

Bothrydiopisalia Scheerpeltz, 1966b: 20

Type species: Leptusa bosnica Eppelsheim, 1892

Bryopisalia Pace, 1989: 239

Type species: Leptusa pilatensis Scheerpeltz, 1972

Chondrelytropisalia Scheerpeltz, 1976c: 55

Type species: Leptusa nepalica Scheerpeltz, 1976

Chondridiopisalia Pace, 1983c: 85

Type species: Leptusa granulifera Pace, 1983

Chondridiopisalia Scheerpeltz, 1966b: 19 [nomen nudum, Newton & Thayer 2003]

Type species: Leptusa granulifera Scheerpeltz, 1966

Chondridiopisalia Lohse, 1974: 53 [ Newton & Thayer 2003]

Chondropisalia Scheerpeltz, 1966b: 14

Type species: Leptusa schaschli Ganglbauer, 1897

Cidnopisalia Pace, 1989: 235

Type species: Leptusa cavallensis Pace, 1980

Comphopisalia Scheerpeltz, 1966b: 16

Type species: Leptusa brucki Scriba, 1867

Craspedopisalia Scheerpeltz, 1966b: 19

Type species: Leptusa carinata Bernhauer, 1909

Cryptopisalia Pace, 1983c: 88

Type species: Leptusa montisparii Scheerpeltz, 1953

Cryptopisalia Pace, 1983e: 129 [ Newton & Thayer 2003]

Cyllopisalia Pace, 1982a: 40

Type species: Leptusa difformis Mulsant & Rey, 1853

Cyllopisalia Pace, 1983c: 79 [ Smetana 2004: 431]

Type species: Leptusa difformis Mulsant & Rey, 1853

Parapisalia Scheerpeltz, 1966b: 18 [synonymized by Pace 1982a: 40]

Type species: Leptusa difformis Mulsant & Rey, 1853

Sipalia Mulsant & Rey, 1853b: 32 View in CoL [synonymized by Pace 1982a: 40]

Type species: Homalota difformis Mulsant & Rey, 1853

Dendroleptusa Pace, 1983c: 65

Type species: Leptusa fumida Erichson, 1839

Drepanoleptusa Pace, 1982d: 93

Type species: Leptusa annapurnensis Pace, 1982

Dysleptusa Pace, 1982c: 580

Type species: Oxypoda fuliginosa Aubé, 1850

Ectinopisalia Scheerpeltz, 1966b: 18

Type species: Leptusa baldensis Ganglbauer, 1895

Emmelopisalia Pace, 1983c: 92

Type species: Leptusa norica Pace, 1979

Entomophallopisalia Pace, 1983c: 90

Type species: Leptusa asturiensis Eppelsheim, 1880

Eospisalia Pace, 1982c: 583

Type species: Leptusa ishizuchiensis Pace, 1982

Eremopisalia Pace, 1989: 196

Type species: Leptusa marianii Pace, 1978

Eucryptusa Casey, 1906: 345

Type species: Silusa nanula Casey, 1893

Evaniopisalia Pace, 1982a: 38

Type species: Leptusa ultracollensis Pace, 1982

Exaeretopisalia Pace, 1989: 233

Type species: Leptusa cansigliensis Pace, 1975

Gallopisalia Pace, 1989: 179

Type species: Leptusa lapidicola Brisout de Barneville, 1863

Gatineauleptusa Klimaszewski in Klimaszewski et al., 2018: 363

Type species: Leptusa gatineauensis Klimaszewski & Pelletier in Klimaszewski et al., 2004

Geopisalia Pace, 1984: 214

Type species: Leptusa peyerimhoffi Bernhauer, 1935

Gnopheropisalia Pace, 1983c: 76

Type species: Leptusa granulipennis Eppelsheim, 1880

Habropisalia Pace, 1989: 216

Type species: Leptusa laticeps Scheerpeltz, 1972

Halmaeusa Kiesenwetter in Kiesenwetter & Kirsch, 1877: 160 [*]

Type species: Halmaeusa antarctica Kiesenwetter, 1877

Antarctophytosus Enderlein, 1909: 377 View in CoL [synonymized by Steel 1964: 366]

Type species: Phytosus atriceps Waterhouse, 1875

Paraphytosus Cameron, 1917c: 125 View in CoL [synonymized by Steel 1964: 366]

Type species: Phytosus atriceps Waterhouse, 1875

Austromalota Brèthes, 1925: 170 View in CoL [synonymized by Steel 1964: 366]

Type species: Austromalota rufimixta Brèthes, 1925

Hemipasilia Pace, 1989: 90

Type species: Leptusa oertzeni Eppelsheim, 1888

Heteroleptusa Pace, 1989: 160

Type species: Sipalia frontalis Casey, 1893

Heterotyphlopasilia Pace, 1983c: 66

Type species: Leptusa serbica Scheerpeltz, 1935

Homopisalia Pace, 1982c: 584

Type species: Leptusa bengalensis Pace, 1982

Ischnopisalia Pace, 1989: 226

Type species: Leptusa poggii Pace, 1978

Kochliodepisalia Pace, 1996a: 35

Type species: Leptusa spirarum Pace, 1996

Koilokoryphepisalia Pace, 1999a: 210

Type species: Leptusa argentierensis Pace, 1999

Lasiopisalia Pace, 1983c: 76

Type species: Leptusa crenulata Bernhauer, 1899

Lathripisalia Pace, 1983e: 135

Type species: Leptusa montispasubii Pace, 1975

Leptopasilia Scheerpeltz, 1966b: 13

Type species: Leptusa hopffgarteni Eppelsheim, 1881

Leptusa Kraatz, 1856b: 60 View in CoL

Type species: Bolitochara pulchella Mannerheim, 1830

Sipalia Mulsant & Rey, 1853a: 45 View in CoL [ Newton & Thayer 2003]

Liopisalia Pace, 1989: 177

Type species: Leptusa serullazi Sainte-Claire Deville, 1919

Megacolypisalia Pace, 1983e: 133

Type species: Leptusa laevicauda Scheerpeltz, 1958

Melopisalia Pace, 1983c: 88

Type species: Leptusa pasubiana Scheerpeltz, 1953

Metaplasmopasilia Pace, 1989: 104

Type species: Leptusa apennina Holdhaus, 1924

Microcolypisalia Pace, 1989: 196

Type species: Leptusa occulta Pace, 1975

Micropisalia Scheerpeltz, 1948: 164

Type species: Leptusa alpestris Scheerpeltz, 1935

Myopopisalia Scheerpeltz, 1966b: 14

Type species: Leptusa bonvouloiri Brisout de Barneville, 1867

Nanoglossa Fauvel, 1868a: 379 [*]

Type species: Microglossa chilensis Fauvel, 1866

Microglossa Fauvel, 1866b: 282 [synonymized by Fauvel 1868a: 379]

Type species: Microglossa chilensis Fauvel, 1866

Microglossa Fauvel, 1866a: 33 [ Newton & Thayer 2003]

Type species: Microglossa chilensis Fauvel, 1866

Pareuryusa Scheerpeltz, 1972d: 131 View in CoL [synonymized by Pace 1989: 110]

Type species: Euryusa parallela Fairmaire & Germain, 1862

Nanopisalia Pace, 1983c: 94

Type species: Homalota piceata Mulsant & Rey, 1853

Micropisalia Scheerpeltz, 1966b: 20 [synonymized by Pace 1983c: 94]

Type species: Homalota piceata Mulsant & Rey, 1853

Nanopisalia Pace, 1983e: 132 [ Newton & Thayer 2003]

Type species: Homalota piceata Mulsant & Rey, 1853

Neopisalia Scheerpeltz, 1966b: 16

Type species: Leptusa microphthalma Reitter, 1887

Nesopisalia Pace, 1992b: 112

Type species: Leptusa centralis Pace, 1992

Stenoleptusa Scheerpeltz, 1966b: 10 [synonymized by Assing 2009c: 1286]

Type species: Bolitochara laeviuscula Hochhuth, 1849

Oligopisalia Scheerpeltz, 1966b: 20

Type species: Leptusa flavicornis Brancsik, 1874

Oncopisalia Pace, 1982c: 589

Type species: Leptusa monachorum Bernhauer, 1912

Oreopisalia Pace, 1983c: 82

Type species: Leptusa hoelzeli Scheerpeltz, 1948

Pachygastropisalia Scheerpeltz, 1966b: 15

Type species: Leptusa lativentris Sharp, 1873

Pachygluta Thomson, 1858: 34

Type species: Oxypoda ruficollis Erichson, 1839

Parapisalia Scheerpeltz, 1948: 159

Type species: Leptusa puellaris Hampe, 1863

Pasilia Mulsant & Rey, 1871: 226

Type species: Homalota nubigena Kiesenwetter, 1861

Pasilia Mulsant & Rey, 1872b: 316 [ Newton & Thayer 2003]

Pasilia Mulsant & Rey, 1873c: 73 [ Newton & Thayer 2003]

Pisalia Mulsant & Rey, 1871: 234

Type species: Homalota globulicollis Mulsant & Rey, 1853

Pisalia Mulsant & Rey, 1872b: 324 [ Newton & Thayer 2003]

Pisalia Mulsant & Rey, 1873c: 73 [ Newton & Thayer 2003]

Tropidiopasilia Scheerpeltz, 1966b: 11 [synonymized by Pace 1989: 190]

Type species: Leptusa tricolor Scriba, 1870

Planepisalia Pace, 1989: 217

Type species: Leptusa kochiana Bernhauer, 1936

Protoleptusa Semenov in Semenov et al., 2001: 156

Type species: Leptusa defuncta Semenov, Perkovsky & Petrenko, 2001

Rhombopisalia Scheerpeltz, 1953: 66

Type species: Leptusa knabli Bernhauer, 1909

Roubaliusa Scheerpeltz, 1966b: 13

Type species: Leptusa storkani Roubal, 1916

Scelopisalia Scheerpeltz, 1966b: 18

Type species: Leptusa gracilipes Krauss, 1899

Scoliophallopisalia Pace, 1989: 232

Type species: Leptusa pascuorum Pace, 1975

Scoliophallopisalia Pace, 1981: 115 [nomen nudum, Newton & Thayer 2003]

Scoliophallopisalia Anonymous, 1984: 249 [ Newton & Thayer 2003]

Type species: Leptusa blesioi Pace, 1981

Scolioplatypisalia Pace, 1989: 86

Type species: Leptusa munelensis Apfelbeck, 1907

Stictopisalia Scheerpeltz, 1966b: 16

Type species: Leptusa eximia Kraatz, 1856

Synpisalia Scheerpeltz, 1966b: 16 [synonymized by Pace 1983c: 56]

Type species: Leptusa caucasica Eppelsheim, 1877

Toxophallopisalia Pace, 1983c: 99

Type species: Leptusa austriaca Scheerpeltz, 1935

Trichopasilia Scheerpeltz, 1966b: 12

Type species: Leptusa major Bernhauer, 1900

Typhlopasilia Ganglbauer, 1895: 278

Type species: Leptusa kaufmanni Ganglbauer, 1895

Ulitusa Casey, 1906: 347

Type species: Ulitusa cribratula Casey, 1906

Yunnaleptusa Assing, 2008b: 256

Type species: Leptusa curvata Assing, 2006

16 species are included in this genus from South America.

Characters/Description: Lynch-Arribálzaga 1884: 87 (characters); Fenyes 1920: 120 (characters), 122 (characters subgenera Typhlosilia and Leptusa ), 123 (characters subgenera Eucryptusa and Pachygluta ), 124 (characters subgenera Oreusa and Pasilia ), 125 (characters subgenera Pisalia ); Pace 1989: 25 (characters), 110 (characters of the subgenus Nanoglossa ), 244 (characters of the subgenus Halmaeusa ).

Key: Bernhauer 1900a: 402 (Subgenera and some species); Fenyes 1920: 121 (subgenera); Pace 1989: 25 (subgenera), 110 (species of the subgenus Nanoglossa ), 246 (species of the subgenus Halmaeusa ); Pace 2001a: 12 (Chilean species).

Distribution.

South America: Chile, Argentina, Falkland Isl.

Other regions: Nearctic, Palaearctic, Ethiopian, Oriental, Australian, Antarctic.

0448. Meoticaphaena Pace, 2015e: 72

Type species: Meotica cordobensis Bernhauer, 1934

Only one species is known from and is restricted to South America.

Distribution.

South America: Argentina.

0449. Phymatura J. Sahlberg, 1876: 85

Type species: Bolitochara brevicollis Kraatz, 1856

Venusa Casey, 1906: 272 View in CoL [synonymized by Ashe 1992: 362]

Type species: Venusa picta Casey, 1906

3 species are included in this genus from South America.

Characters/Description: Fenyes 1920: 116 (characters); Ashe 1992: 366 (characters in key), 371 (characters). Distribution.

South America: Brazil, Argentina.

Other regions: Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian.

0450. Prosoponotha Pace, 1999c: 146

Type species: Prosoponotha chilensis Pace, 1999

5 species are known from and are restricted to South America.

Distribution.

South America: Brazil, Chile.

0451. Tanystetha Pace, 2002: 137

Type species: Tanystetha brasiliensis Pace, 2002

Only one species is known from and is restricted to South America.

Distribution.

South America: Brazil. Subtribe Dinardopsina Bernhauer & Scheerpeltz 1926

0452. Dinardopsis Bruch, 1917: 257 ( Fig. 52 View FIGURES 52–63 )

Type species: Dinardopsis solenopsidicola Bruch, 1917

2 species are known from and are restricted to South America.

Distribution.

South America: Brazil, Argentina.

References

Anonymous (1984) Insecta: Part B, Coleoptera. Zoological Record, 118 (13 B), 1 - 362.

Ashe, J. S. (1992) Phylogeny and revision of genera of the subtribe Bolitocharina (Coleoptera: Staphylinidae: Aleocharinae). University of Kansas Science Bulletin, 54, 335 - 406. https: // doi. org / 10.5962 / bhl. part. 19642

Assing, V. (2008 b) Seven new species and additional records of Palaearctic Leptusa, primarily from Yunnan, China (Coleoptera: Staphylinidae: Aleocharinae). Linzer biologische Beitrage, 40 (1), 251 - 273.

Assing, V. (2009 c) New species, new synonymies, and additional records of Leptusa from Turkey and Iran (Coleoptera: Staphylinidae: Aleocharinae). Linzer biologische Beitrage, 41 (2), 1285 - 1305.

Bernhauer, M. (1899) Dritte Folge neuer Staphyliniden aus Europa, nebst synonymischen und anderen Bemerkungen. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 49, 15 - 27.

Bernhauer, M. (1900 a) Die Staphyliniden-Gattung Leptusa Kraatz, nebst einer analytischen Bestimmungstabelle der palaarktischen Arten. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 50, 399 - 432.

Bernhauer, M. & Scheerpeltz, O. (1926) Staphylinidae VI. In: Schenkling, S. (Ed.), Coleopterorum Catalogus, vol. 5, pars 82. W. Junk, Berlin, pp. 499 - 988.

Bierig, A. (1937 b) Algunos nuevos estafilinidos (Col. Staph.) Cubanos. (20 ª. contribucion al conocimiento de los estafilinidos). Memorias de la Sociedad Cubana de Historia Natural Felipe Poey , 11 (4), 273 - 283.

Blackwelder, R. E. (1952) The generic names of the beetle family Staphylinidae, with an essay on genotypy. Bulletin of the United States National Museum, 200, iv + 483 pp.

Brethes, J. (1925) Un Coleoptere et un Diptere nouveaux de la Georgie du Sud. Comunicaciones del Museo Nacional de Historia Natural Bernardino Rivadavia , 2, 169 - 173.

Bruch, C. (1917) Descripcion de un nuevo genero y de dos nuevas especies de estafilinidos mirmecofilos. Anales de la Sociedad Cientifica Argentina, 82, 257 - 264. [Note: nominal year of publication 1916]

Cameron, M. (1917 c) Description of a new genus of Staphylinidae. The Entomologist's Monthly Magazine, 53, 125.

Casey, T. L. (1893) Coleopterological notices, V. Annals of the New York Academy of Sciences, 7, 281 - 601, pl. 1. [Note: nominal year of publication 1894]

Casey, T. L. (1906) Observations on the staphylinid groups Aleocharinae and Xantholinini chiefly of America. Transactions of the Academy of Science of St. Louis, 16 (6), 125 - 434.

Coiffait, H. & Saiz, F. (1967) Aleocharidae du Chili. I. - Tribus Oligotini, Myllaenini, Bolitocharini (Col. Staphylinoidea). Bulletin de la Societe d'Histoire Naturelle de Toulouse, 103, 51 - 98.

Enderlein, G. (1909) Die Insekten des Antarktischen Gebietes. In: Deutsche Sudpolar-Expedition 1901 - 1903. Im Auftrage des Reichsamtes des Innern herausgegeben von Erich von Drygalski, Leiter der Expedition. Vol. 10, Zoologie, Vol. 2. Georg Reimer, Berlin. pp. 361 - 528, pls. 40 - 63. [Note: nominal year of publication 1908]

Fairmaire, L. & Germain, P. (1862) Revision des Coleopteres du Chili (suite). Annales de la Societe Entomologique de France, 4 (1), 405 - 456. [Note: nominal year of publication 1861]

Fauvel, A. (1866 b) Faune du Chili. Insectes Coleopteres. Staphylinides. Bulletin de la Societe Linneenne de Normandie, 10, 250 - 353, pl. 4. [Note: nominal year of publication 1864 - 1865]

Fauvel, A. (1866 a) Faune du Chili. Insectes Coleopteres. Staphylinides. Notices Entomologiques, 4 (1), 1 - 103, pl. 4.

Fauvel, A. (1868 a) [Untitled]. Bulletin de la Societe Linneenne de Normandie, 2 (2), 379. [Note: nominal year of publication 1867]

Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae. Subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum. Fascicule 173 b. L. Desmet-Verteneuil, Brussels, pp. 111 - 414.

Ganglbauer, L. (1895) Die Kafer von Mitteleuropa. Die Kafer der osterreichisch-ungarischen Monarchie, Deutschlands, der Schweiz, sowie des franzosischen und italienischen Alpengebietes. Vol. 2, Familienreihe Staphylinoidea, I. Theil: Staphylinidae, Pselaphidae. Carl Gerold's Sohn, Vienna. vi + 881 pp.

Gravenhorst, J. L. C. (1806) Monographia Coleopterorum Micropterorum. H. Dieterich, Gottingae [Gottingen]. xvi + 248 pp., 1 pl.

Hampe, C. (1863) Ein kleiner Beitrag zur gross-osterreichischen Kaferfauna. Wiener Entomologische Monatschrift, 7 (8), 285 - 290.

Kiesenwetter, E. A. H. von & Kirsch, T. (1877) Die Kaferfauna der Auckland-Inseln, nach Herm. Krone's Sammlungen beschrieben. Deutsche Entomologische Zeitschrift, 21 (1), 153 - 174.

Kraatz, G. (1856 b) Naturgeschichte der Insecten Deutschlands. Abteilung 1, Coleoptera, vol. 2, Staphylinii. Lief. 1 - 2. Nicolaische Buchhandlung, Berlin, vii + 1 - 376 pp. [Note: nominal year of publication 1858]

Lohse, G. A. (1974) Hypocyphtinae, Aleocharinae [in part], In: Freude, H., Harde, K. W. & Lohse, G. A. (Eds.). Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae), Pselaphidae. Goecke & Evers, Krefeld. pp. 7 - 72, 220 - 292.

Lynch-Arribalzaga, F. (1884) Estafilinos de Buenos Aires. Boletin de la Academia Nacional de Ciencias en Cordoba, 7 (1 - 3), 5 - 392.

Mannerheim, C. G. (1830) Precis d'un nouvel arrangement de la famille des brachelytres, de l'ordre des insectes coleopteres. St. Petersbourg. 87 pp.

Mulsant, E. & Rey, C. (1853 b) Description de quelques Coleopteres nouveaux ou peu connus, de la tribu des Brachelytres. Annales de la Societe Linneenne de Lyon (n. s.), 1, 22 - 72, pl. 2.

Mulsant, E. & Rey, C. (1853 a) Description de quelques Coleopteres nouveaux ou peu connus, de la tribu des Brachelytres. Opuscules Entomologiques, 2, 35 - 85, pl. 2. [Note: nominal year of publication 1852]

Mulsant, E. & Rey, C. (1871) Histoire Naturelle des Coleopteres de France. Brevipennes. Aleochariens [Huitieme Branche, Bolitocharaires]. Deyrolle Fils, Paris. 321 pp., pls. 1 - 5.

Mulsant, E. & Rey, C. (1872 b) Tribe des brevipennes: Famille des aleochariens: Huitieme branche: Bolitocharaires. Annales de la Societe Linneenne de Lyon (n. s.), 19, 91 - 413, 426, pl. 1 - 5.

Mulsant, E. & Rey, C. (1873 c) Tribe des Coleopteres brevipennes: Famille des aleochariens: Premiere branche: Bolitocharaires. Memoires de l'Academie des Sciences, Belles-Lettres et Arts de Lyon, Classe des Sciences, 2 (19), 73 - 123. [Note: nominal year of publication 1871 - 1872]

Newton, A. F. & Thayer, M. K. (2003) Catalog of higher taxa, genera and subgenera of Staphyliniformia [online]. Chicago: Field Museum of Natural History [last update 03 November 2005]. Available from URL: <http: // fm 1. fieldmuseum. org / peet /> (accessed November 2016).

Pace, R. (1981) Nuovo contributo alla conoscenza delle specie bresciane del genere Leptusa Kraatz (Coleoptera, Staphylinidae). Natura Bresciana, 17, 114 - 123. [Note: nominal year of publication 1980]

Pace, R. (1982 c) Leptusa raccolte dal Dr. Claude Besuchet nella Penisola Iberica, nella regione balcanica, in Turchia e in estremo oriente (Coleoptera, Staphylinidae). Revue Suisse de Zoologie, 89, 579 - 594. https: // doi. org / 10.5962 / bhl. part. 82461

Pace, R. (1982 a) Nuovo contributo alla conoscenza delle specie italiane del genere Leptusa Kraatz (Coleoptera Staphylinidae). Bollettino della Societa Entomologica Italiana, 114 (1 - 3), 34 - 41.

Pace, R. (1982 d) Aleocharinae del Nepal e dell'India settentrionale raccolte dal Prof. Herbert Franz. I. Bolitocharini (Coleoptera Staphylinidae). Bollettino della Societa Entomologica Italiana, 114 (4 - 7), 87 - 96.

Pace, R. (1983 c) Risultati dello studio delle specie del genere Leptusa Kraatz della collezione Scheerpeltz al Naturhistorisches Museum di Vienna (Coleoptera, Staphylinidae) (XXII Contributo alla conoscenza delle Aleocharinae). Annalen des Naturhistorischen Museums in Wien, 85 B, 53 - 102.

Pace, R. (1983 e) Le specie del genere Leptusa del Monte Baldo (Coleoptera, Staphylinidae). Bollettino del Museo Civico di Storia Naturale di Verona, 9, 121 - 138. [Note: nominal year of publication 1982]

Pace, R. (1984) Nuove Aleocharinae microftalme Mediterranee e dell'Iran, del Museum d'Histoire Naturelle di Ginevra (Coleoptera Staphylinidae). Archives des Sciences (Geneve), 37, 211 - 219.

Pace, R. (1989) Monografia del genere Leptusa Kraatz (Coleoptera, Staphylinidae). Memorie del Museo Civico di Storia Naturale di Verona (A: Biologica), 2 (8), 1 - 307.

Pace, R. (1992 b) Il genere Leptusa Kraatz della sottoregione indocinese (Taiwan e Vietnam). Monografia del genere Leptusa Kraatz, Supplemento I (Coleoptera: Staphylinidae). Elytron, 5, 111 - 119. [Note: nominal year of publication 1991]

Pace, R. (1995) Nuove specie di Leptusa Kr. di Taiwan. Monografia del genere Leptusa Kraatz: Supplemento V (Coleoptera, Staphylinidae). Bollettino della Societa Entomologica Italiana, 126 (3), 243 - 248. [Note: nominal year of publication 1994]

Pace, R. (1996 a) Nuove specie di Leptusa Kraatz di Taiwan. Monografia del genere Leptusa Kraatz, supplemento VI (Coleoptera Staphylinidae). Bollettino della Societa Entomologica Italiana, 128 (1), 29 - 36.

Pace, R. (1997 a) Specie del genere Leptusa in Cina. Monografia del genere Leptusa Kraatz: Supplemento VII (Coleoptera, Staphylinidae). Revue Suisse de Zoologie, 104 (4), 751 - 760. https: // doi. org / 10.5962 / bhl. part. 80018

Pace, R. (1999 a) Nuove specie del genere Leptusa Kraatz raccolte da Manfred Kahlen (Monografia del genere Leptusa Kraatz: Supplemento VIII) (Coleoptera, Staphylinidae). Veroffentlichungen des Tiroler Landesmuseums Ferdinandeum, 79, 207 - 214.

Pace, R. (1999 c) Aleocharinae del Cile (Coleoptera, Staphylinidae), (114 º Contributo alla conoscenza delle Aleocharinae). Bollettino del Museo Civico di Storia Naturale di Verona, 23, 119 - 210.

Pace, R. (2001 a) Nuove specie Cilene del genere Leptusa. Monografia del genere Leptusa Kraatz: Supplemento XII (162 º Contributo alla conoscenza delle Aleocharinae). Lavori della Societa Veneziana di Scienze Naturali, 26, 7 - 14.

Pace, R. (2002) Gyrophaenini e Bolitocharini del Sudamerica (Coleoptera, Staphylinidae). Nouvelle Revue d'Entomologie (n. s.), 19 (2), 115 - 139.

Pace, R. (2007) Nuovi dati faunistici e tassonomici su specie di Aleocharinae del Sudamerica delle tribu Gyrophaenini, Placusini, Diestotini, Homalotini, Bolitocharini, Paglini e Falagriini (Coleoptera, Staphylinidae). Beitrage zur Entomologie, 57 (1), 211 - 239. https: // doi. org / 10.21248 / contrib. entomol. 57.1.211 - 239

Pace, R. (2015 e) Biodiversity of Aleocharinae from Andean and Brazilian sub-regions: new data, new genera and new species (Coleoptera Staphylinidae). In: Memoirs on Biodiversity vol. 3 (Biodiversity of South America II), WBA Project, Verona, 3, pp. 39 - 173.

Sahlberg, J. (1876) Enumeratio coleopterorum brachelytrorum Fenniae. Systematisk forteckning ofver de inom Finlands naturalhistoriska omrade hittills funna Coleoptera Brachelytra ... I. Staphylinidae. Acta Societatis pro Fauna et Flora Fennica, 1 (3), 1 - 248.

Scheerpeltz, O. (1953) Rhombopisalia, eine neue Untergattung der Grossgattung Leptusa Kr., mit einer Bestimmungstabelle der bis jetzt aus dieser Untergattung bekannt gewordenen Arten (Col. Staphylinidae). (52. Beitrag zur Kenntnis der palaarktischen Staphyliniden). Studi Trentini di Scienze Naturali, 30 (1), 65 - 77.

Scheerpeltz, O. (1958) Wissenschaftliche Ergebnisse der von Herrn Dr. K. Lindberg, Lund, im Jahre 1956 nach der Turkei und Armenien unternommenen Reise. Coleoptera - Staphylinidae. (82. Beitrag zur Kenntnis der palaarktischen Staphyliniden). Entomologisk Tidskrift (supplement), 78 (10), 3 - 37. [Note: nominal year of publication 1957]

Scheerpeltz, O. (1966 b) Die neue Systematik der Grossgattung Leptusa Kraatz (Col. Staphylinidae). (133. Beitrag zur Kenntnis der palaarktischen Staphyliniden). Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 105 / 106, 5 - 55.

Scheerpeltz, O. (1972 d) Wissenschaftliche Ergebnisse der Studienreise von Gy. Topal nach Sudwest-Argentinien (Coleoptera: Staphylinidae). Folia Entomologica Hungarica (s. n.), 25 (Suppl.), 1 - 268, pl. 5.

Scheerpeltz, O. (1976 c) Wissenschaftliche Ergebnisse der von Prof. Dr. H. Janetschek im Jahre 1961 in das Mt. - Everest-Gebiet Nepals unternommenen Studienreise (Col. Staphylinidae). (157. Beitrag zur Kenntnis der palaartischen Staphyliniden, gleichzeitig 12. Beitrag zur Kenntnis der orientalischen Staphyliniden). Khumbu Himal, Ergebnisse des Forschungsunternehmens Nepal Himalaya, 5, 3 - 75.

Seevers, C. H. (1978) A generic and tribal revisions of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana Zoloogy, 71, vi + 289 pp.

Semenov, V. B., Perkovsky, E. E. & Petrenko, A. A. (2001) Pervaya nakhodka aleokharin (Coleoptera, Staphylinidae, Aleocharinae) iz rovenskogo yantarya. Reports of the National Academy of Sciences of Ukraine, 2001 (7), 155 - 158.

Smetana, A. (2004) Subfamily Aleocharinae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Volume 2. Hydrophiloidea - Histeroidea - Staphylinoidea. Apollo Books, Stenstrup, pp. 353 - 494.

Steel, W. O. (1964) Insect of Campbell Island. Coleoptera: Staphylinidae. Pacific Insects Monograph, 7, 340 - 375.

Thomson, C. G. (1858) Forsok till uppstallning af Sveriges Staphyliner. Ofversigt af Kongliga Vetenskaps-Akademiens Forhandlingar, 15, 27 - 40.

Thomson, C. G. (1859) Skandinaviens Coleoptera, synoptiskt bearbetade. Vol. I. Berlingska Boktryckeriet, Lund, 290 pp.

Figures

FIGURES 52–63. Habitus; Aleocharinae (52–63); 52, after Bruch 1917; 53, Peru: department of Madre de Dios (CEMT); 54, Peru: department of Madre de Dios (CEMT); 55, after Pace 2015e; 56, after Pace 2015a; 57, after Pace 2011b; 58, after Pace 2007; 59, after Pace 2008d; 60, after Pace 2015e; 61, after Pace 2014c; 62, after Pace 2011b; 63, after Pace 2011b; slightly modified. Scale bars: 1 mm.