Carineta bilineosa Walker, 1858b, 2019

Sanborn, Allen F., 2019, The cicadas (Hemiptera: Cicadidae) of Bolivia including the descriptions of fifteen new species, the resurrection of one genus and two species, seven new combinations, six new synonymies, and twenty-eight new records, Zootaxa 4655 (1), pp. 1-104 : 55-57

publication ID

https://doi.org/ 10.11646/zootaxa.4655.1.1

publication LSID

lsid:zoobank.org:pub:3B65A3A8-2D1E-4031-8BD4-5A1A327C4ADE

DOI

https://doi.org/10.5281/zenodo.4439589

persistent identifier

https://treatment.plazi.org/id/039787CA-595A-FFF7-FF51-FE82361FCFDB

treatment provided by

Plazi

scientific name

Carineta bilineosa Walker, 1858b
status

stat. nov.

Carineta bilineosa Walker, 1858b View in CoL rev. stat., new record

( Fig. 8 View FIGURE 8 )

Carineta bilineosa Walker 1858b: 12 View in CoL . ( Brazil) rev. stat.

Carineta obtusa Walker 1858b: 14 View in CoL . ( Brazil) rev. stat., n. syn.

Carineta tenuistriga Walker 1858c: 311 View in CoL . (Tejuca, Rio de Janeiro, Brazil) rev. stat., n. syn.

Carineta diplographa Berg 1879: 212 View in CoL . (Córpus, Misiones, Argentina) rev. stat., n. syn.

Carineta fasciculata View in CoL (non Germar) Berg 1883: 184.

Carineta fasciculata View in CoL ? (non Germar) d’Utra 1908: 354–355, Fig 2 View FIGURE 2 .

Carineta fasciculata View in CoL (non Germar) Hempel 1913a: 94.

Carineta fasciculata View in CoL (non Germar) Hempel 1913b: 139.

Carineta fasciculata View in CoL (non Germar) Hempel 1913c: 197.

Carineta fasciculata View in CoL (non Germar) Hempel 1913d: 3.

Carineta fasciculata View in CoL (non Germar) Delétang 1919: 64.

Carinetta (sic) fasciculata View in CoL (non Germar) Moreira 1921: 125.

Carineta fasciculata View in CoL (non Germar) Costa Lima 1922a: 111.

Carineta fasciculata View in CoL (non Germar) Costa Lima 1922b: 113.

Carineta fasciculata View in CoL (non Germar) Moreira 1928: 24.

Carineta fasciculata View in CoL (non Germar) Moreira 1929: 139–140.

Carineta fasciculata View in CoL (non Germar) Fonseca & Autuori 1932: 60, Figs 29A–29C.

Carineta fasciculata View in CoL (non Germar) Monte 1932: 25, Fig 19 View FIGURE 19 .

Carineta fasciculata View in CoL (non Germar) Fonseca 1934: 267.

Carineta fasciculata View in CoL (non Germar) Wymore1934: 885.

Carineta fasciculata View in CoL (non Germar) Costa Lima 1936: 131.

Carineta fasciculata View in CoL (non Germar) Fonseca & Araujo 1939: 285, Plate XII, Fig 5 View FIGURE 5 .

Carineta fasciculata View in CoL (non Germar) Costa Lima 1942: 11, Fig 5 View FIGURE 5 .

Fidicina fasciculata View in CoL (non Germar) Fonseca 1945: 297.

Carineta fasciculata View in CoL (non Germar) Fonseca 1945: 301.

Carineta fasciculata View in CoL (non Germar) Torres 1945: 4, 8–9, Fig 2 View FIGURE 2 .

Carineta fasciculata View in CoL (non Germar) Torres 1946: 10, Plate 2, Fig 4 View FIGURE 4 .

Carineta fasciculata View in CoL (non Germar) Torres 1948b: 73, Figs 1A View FIGURE 1 , 2A View FIGURE 2 .

Carineta fasciculata View in CoL (non Germar) Pesson 1951: 1502.

Carineta fasciculata View in CoL (non Germar) Torres & China 1955: 479–480 View Cited Treatment , Figs 1 View FIGURE 1 a–1b.

Carineta fasciculata View in CoL (non Germar) Vayssière 1955: 254.

Carineta fasciculata View in CoL (non Germar) Müller 1956: 192.

Carineta fasciculata View in CoL (non Germar) Gomez-Menor Ortega 1957: 28.

Fidicina fasciculata View in CoL (non Germar) Mariconi 1958: 331–332.

Carineta fasciculata View in CoL (non Germar) Hayward 1960: 35.

Carineta fasciculata View in CoL (non Germar) Torres 1963b: 50, 52, 55, 56, Figs. 14–15 View FIGURE 14 View FIGURE 15 .

Carineta fasciculata View in CoL (non Germar) Heinrich & Pupin Neto 1964: 5.

Carineta fasciculata View in CoL (non Germar) Heinrich 1967: 43.

Carineta fasciculata View in CoL (non Germar) Lekić 1967: 3.

Carineta fasciculata View in CoL (non Germar) Le Pelley 1968: 306, 467.

Carineta fasciculata View in CoL (non Germar) Weidner & Wagner 1968: 142.

Carineta fasciculata View in CoL (non Germar) Martinelli & Zucchi 1987a: 469.

Carineta fasciculata View in CoL (non Germar) Martinelli & Zucchi 1987b: 470.

Carineta fasciculata View in CoL (non Germar) Santos 1988: 27.

Carineta fasciculata View in CoL (non Germar) Martinelli & Zucchi 1989b: 13–16, 18, 20 View Cited Treatment , Fig 1 View FIGURE 1 .

Carineta fasciculata View in CoL (non Germar) Martinelli 1990: 13.

Carineta fasciculata View in CoL (non Germar) Martinelli & Zucchi 1997a: 208 View Cited Treatment .

Carineta fasciculata View in CoL (non Germar) Martinelli & Zucchi 1997b: 135–141 View Cited Treatment , Figs 1–2 View FIGURE 1 View FIGURE 2 , 14 View FIGURE 14 , 18 View FIGURE 18 , Table 1.

Carineta fasciculata View in CoL (non Germar) Albuquerque, Martinelli, Ros & Stülp 2000: 196.

Carineta fasciculata View in CoL (non Germar) Fornazier & Martinelli 2000: 1176.

Carineta fasciculata View in CoL (non Germar) Fornazier & Rocha 2000: 1167, 1169.

Carineta fasciculata View in CoL (non Germar) Scarpellini 2001: 66.

Carineta fasciculata View in CoL (non Germar) Reis, Souza & Venzon 2002: 84.

Carineta fasciculata View in CoL (non Germar) Almeida 2004: 104–105.

Carineta fasciculata View in CoL (non Germar) Martinelli 2004: 518–521 View Cited Treatment , 525–526, 529, Figs 18.1–18.2, 18.4d, 18.4g.

Carineta fasciculata View in CoL (non Germar) Martinelli, Maccagnan, Ribeiro, Pereira & Santos 2004: 619 View Cited Treatment .

Carineta fasciculata View in CoL (non Germar) Santos & Martinelli 2004: 630.

Carineta fasciculata View in CoL (non Germar) Bolcatto, Medrano & De Santis 2006: 7, 10, Map 1.

Carineta fasciculata View in CoL (non Germar) Martinelli 2006: 1.

Carineta fasciculata View in CoL (non Germar) De Santis, Medrano, Sanborn & Bolcatto 2007: 11–12, Fig 2 View FIGURE 2 .

Carineta fasciculata View in CoL (non Germar) Santos & Martinelli 2007: 311.

Carineta fasciculata View in CoL (non Germar) Lara, Perioto & de Freitas 2010: 116.

Carineta fasciculata View in CoL (non Germar) Maccagnan & Martinelli 2011: 446–448 View Cited Treatment , 450–451, Figs 1 View FIGURE 1 i–1 j, 2f, 3f, 4e.

Carineta fasciculate (sic) (non Germar) Maccagnan & Martinelli 2011: 448, Table 1.

Carineta fasciculata View in CoL (non Germar) Decaro Júnior, Martinelli, Maccagnan & Ribeiro 2012: 1.

Carineta fasciculata View in CoL (non Germar) Sanborn & Heath 2014: 69 View Cited Treatment .

REMARKS. The application of the taxa C. fasciculata , its current synonyms and C. maculosa has been confused historically. However, an image of the Germar type (or at least what Germar considered to be C. fasciculata stored in his collection) located on the website (http://zoomus.lviv.ua/en/germar_collection/) of the Zoological Museum, Ivan Franko National University of Lviv, L’viv, Ukraine (ZMD) has allowed for a definitive distinction of the various taxa. This specimen is from the Germar collection and verifies what Germar considered to be his taxon permitting the clarification and application of the taxon in the literature.

Berg (1883) appears to be the first to synonymize C. fasciculata with Carineta bilineosa rev. stat., Carineta obtusa rev. stat., n. syn., Carineta tenuistriga rev. stat., n. syn., and Carineta diplographa rev. stat., n. syn. Distant (1883b) listed the species as synonyms but with question marks suggesting he was not certain of this synonymy. Distant (1906a) then removed the question marks and considered the species to be synonyms in his catalogue. This interpretation was continued in the later Cicadoidea catalogues ( Metcalf 1963c; Duffels & van der Laan 1985; Sanborn 2013) along with the majority of researches publishing on the species as outlined above.

Jacobi (1907) made several errors in his interpretation of taxa and the concept of each taxon is updated here to clarify the actual taxa. Jacobi (1907) incorrectly identified and illustrated ( Jacobi 1907, Fig. 21) a marked specimen as C. bilineosa (actually C. pilifera Walker, 1858c based on the ovoid body, the longitudinal piceous mark on the posterior dorsal prothorax, the lack of a transverse piceous mark in the ambient fissure, the marks surrounding the mesothoracic sigillae, the presence of the a piceous mark between the anterior arms of the cruciform elevation, greater pilosity, the angle of the junction of the median vein and radius posterior at the node, the shape of the ulnar and apical cells of the fore wing, the dark markings on the female dorsolateral abdominal segment 9, and known distribution in both Bolivia and Ecuador all of which differ from C. bilineosa ). He also identified what would become C. maculosa as C. fasciculata ( Jacobi 1907, Fig. 25). Finally, Jacobi (1951) made comparative references to the similar species C. postica and C. fasciculata (actually C. maculosa ) in the description of C. titschacki Jacobi, 1951 , both of which differ significantly from C. bilineosa so that his concept of C. bilineosa is incorrect as is his concept of C. fasciculata which was later to be determined as a new species ( Torres 1948a).

Torres (1948a) started to clarify the taxa by identifying the species illustrated as C. fasciculata in Jacobi (1907) as his new species C. maculosa Torres, 1948a . Torres (1948a) further applied his new taxon to identifications of examples of northern Carineta species previously identified as C. fasciculata . The image of the specimen from the Germar collection has reduced thoracic marking and has the more stout body similar to C. maculosa rather than the heavily marked, thinner body of the current junior synonyms of C. fasciculata . The availability of the image of the Germar specimen compared to the type material of the Walker species in the BMNH and the Berg species in the Museo de La Plata shows that these are not the same taxa and the previously synonymized species Carineta bilineosa rev. stat., Carineta obtusa rev. stat., n. syn., Carineta tenuistriga rev. stat., n. syn., and Carineta diplographa rev. stat., n. syn. are resurrected from junior synonymy with C. fasciculata here although the resurrected species do all represent the same taxon.

The specimen illustrated here ( Fig. 8 View FIGURE 8 ) is from near the type locality of Carineta diplographa rev. stat., n. syn. in Misiones Province, Argentina and matches the image of the holotype in Marino de Remes Lenicov et al. (2015). It has the same narrow body, piceous markings, shape of fore wing cells, position of fore wing veins, shape of timbal cavity as the holotype of C. bilineosa rev. stat. in the BMNH. The specimen also matches the coloration and descriptions of Carineta bilineosa rev. stat., Carineta obtusa rev. stat., n. syn., Carineta tenuistriga rev. stat., n. syn., and Carineta diplographa rev. stat., n. syn. In addition, Berg (1879) distinguishes his new species from C. fasciculata by a list of piceous markings that are missing in C. fasciculata , differences confirmed by the image of the Germar specimen. These data and images confirm that Carineta bilineosa rev. stat., Carineta obtusa rev. stat., n. syn., Carineta tenuistriga rev. stat., n. syn., and Carineta diplographa rev. stat., n. syn. are not synonyms of C. fasciculata and are here resurrected. In addition, the synonymy of Carineta obtusa rev. stat., n. syn., Carineta tenuistriga rev. stat., n. syn., and Carineta diplographa rev. stat., n. syn. with Carineta bilineosa rev. stat. is also confirmed by examination of the Walker type specimens in the BMNH and the image of the holotype of Carineta diplographa rev. stat., n. syn. in Marino de Remes Lenicov et al. (2015).

Martinelli & Zucchi (1997b) illustrate the distribution of C. bilineosa rev. stat. (as C. fasciculata as described above) inhabiting grassland, savannah, open forest habitats, xeric shrub lands and agricultural areas (as defined by Leeuwen et al. 2013) of eastern and southern Brazil, northeastern Argentina, eastern Paraguay and northeastern Bolivia ( Martinelli & Zucchi 1997b). This corresponds with the description of C. diplographa rev. stat., n. syn. described from most northeastern Province of Argentina but the distribution is more expansive than that of the true C. fasciculata (see below) with overlap in the distribution of the two species only in Misiones, Argentina, and the state of and São Paulo in Brazil.

The most similar species in Bolivia is C. cingenda Distant, 1883b . This species differs in the transverse piceous fascia in the ambient fissure of the pronotum and reduced piceous markings of the mesothorax.

DISTRIBUTION. The species has been reported from Argentina, Bolivia, Brazil, and Paraguay as C. fasciculata ( Metcalf 1963c; Duffels & van der Laan 1985; Maccagnan & Martinelli 2011; Sanborn 2011b; 2013; 2018c). However, C. bilineosa rev. stat. and its synonyms have been reported previously only from Argentina and Brazil ( Metcalf 1963c) so this is the first references to C. bilineosa rev. stat. in both Bolivia and Paraguay. The specimen listed by Jacobi (1907) from Bolivia is considered C. pilifera as outlined above.

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Hemiptera

Family

Cicadidae

Genus

Carineta

Loc

Carineta bilineosa Walker, 1858b

Sanborn, Allen F. 2019
2019
Loc

Carineta fasciculata

Sanborn, A. F. & Heath, M. S. 2014: 69
2014
Loc

Carineta fasciculata

Decaro Junior, S. T. & Martinelli, N. & Maccagnan, D. H. B. & Ribeiro, E. S. D. B. P. 2012: 1
2012
Loc

Carineta fasciculate

Maccagnan, D. H. B. & Martinelli, N. M. 2011: 448
2011
Loc

Carineta fasciculata

Lara, R. I. R. & Perioto, N. W. & de Freitas, S. 2010: 116
2010
Loc

Carineta fasciculata

De Santis, Medrano, Sanborn & Bolcatto 2007: 11-12
2007
Loc

Carineta fasciculata

Santos, R. S. & Martinelli, N. M. 2007: 311
2007
Loc

Carineta fasciculata

Bolcatto, P. G. & Medrano, M. C. & De Santis, C. 2006: 7
2006
Loc

Carineta fasciculata

Martinelli, N. M. 2006: 1
2006
Loc

Carineta fasciculata

Martinelli, N. M. 2004: 518-521
2004
Loc

Carineta fasciculata

Santos, R. S. & Martinelli, N. M. 2004: 630
2004
Loc

Carineta fasciculata

Reis, P. R. & Souza, J. C. de & Venzon, M. 2002: 84
2002
Loc

Carineta fasciculata

Scarpellini, J. R. 2001: 66
2001
Loc

Carineta fasciculata

Albuquerque, F. A. & Martinelli, N. M. & Ros, A. B. & Stulp, M. 2000: 196
2000
Loc

Carineta fasciculata

Fornazier, M. J. & Martinelli, N. M. 2000: 1176
2000
Loc

Carineta fasciculata

Fornazier, M. J. & Rocha, A. C. da 2000: 1167
2000
Loc

Carineta fasciculata

Martinelli & Zucchi 1997: 135-141
1997
Loc

Carineta fasciculata

Martinelli, N. M. 1990: 13
1990
Loc

Carineta fasciculata

Martinelli & Zucchi 1989: 13-16
1989
Loc

Carineta fasciculata

Santos, B. B. dos 1988: 27
1988
Loc

Carineta fasciculata

Martinelli, N. M. & Zucchi, R. A. 1987: 469
1987
Loc

Carineta fasciculata

Martinelli, N. M. & Zucchi, R. A. 1987: 470
1987
Loc

Carineta fasciculata

Le Pelley, R. H. 1968: 306
1968
Loc

Carineta fasciculata

Weidner, H. & Wagner, W. 1968: 142
1968
Loc

Carineta fasciculata

Heinrich, W. O. 1967: 43
1967
Loc

Carineta fasciculata

Lekic, M. 1967: 3
1967
Loc

Carineta fasciculata

Heinrich, W. O. & Pupin Neto, J. 1964: 5
1964
Loc

Carineta fasciculata

Torres, B. A. 1963: 50
1963
Loc

Carineta fasciculata

Hayward, K. J. 1960: 35
1960
Loc

Fidicina fasciculata

Mariconi 1958: 331-332
1958
Loc

Carineta fasciculata

Gomez-Menor Ortega, J. 1957: 28
1957
Loc

Carineta fasciculata

Muller, H. J. 1956: 192
1956
Loc

Carineta fasciculata

Torres & China 1955: 479-480
1955
Loc

Carineta fasciculata

Vayssiere, P. 1955: 254
1955
Loc

Carineta fasciculata

Pesson, P. 1951: 1502
1951
Loc

Carineta fasciculata

Torres, B. A. 1948: 73
1948
Loc

Carineta fasciculata

Torres, B. A. 1946: 10
1946
Loc

Fidicina fasciculata

Fonseca, J. P. 1945: 297
1945
Loc

Carineta fasciculata

Fonseca, J. P. 1945: 301
1945
Loc

Carineta fasciculata

Torres, B. A. 1945: 4
1945
Loc

Carineta fasciculata

Costa Lima, A. M. da 1942: 11
1942
Loc

Carineta fasciculata

Fonseca, J. P. & Araujo, R. L. 1939: 285
1939
Loc

Carineta fasciculata

Costa Lima, A. M. 1936: 131
1936
Loc

Carineta fasciculata

Fonseca, J. P. 1934: 267
1934
Loc

Carineta fasciculata

Wymore, F. H. 1934: 885
1934
Loc

Carineta fasciculata

Fonseca & Autuori 1932: 60
1932
Loc

Carineta fasciculata

Monte, O. 1932: 25
1932
Loc

Carineta fasciculata

Moreira, C. 1929: 139-140
1929
Loc

Carineta fasciculata

Moreira, C. 1928: 24
1928
Loc

Carineta fasciculata

Costa Lima, A. M. da 1922: 111
1922
Loc

Carineta fasciculata

Costa Lima, A. da 1922: 113
1922
Loc

Carinetta (sic) fasciculata

Moreira, C. 1921: 125
1921
Loc

Carineta fasciculata

Deletang, L. F. 1919: 64
1919
Loc

Carineta fasciculata

Hempel, A. 1913: 94
1913
Loc

Carineta fasciculata

Hempel, A. 1913: 139
1913
Loc

Carineta fasciculata

Hempel, A. 1913: 197
1913
Loc

Carineta fasciculata

Hempel, A. 1913: 3
1913
Loc

Carineta fasciculata

d’Utra 1908: 354-355
1908
Loc

Carineta fasciculata

Berg, C. 1883: 184
1883
Loc

Carineta diplographa

Berg, C. 1879: 212
1879
Loc

Carineta bilineosa

Walker, F. 1858: 12
1858
Loc

Carineta obtusa

Walker, F. 1858: 14
1858
Loc

Carineta tenuistriga

Walker, F. 1858: 311
1858
Loc

Carineta fasciculata

Almeida 2004: 104–105
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