Loneura javierensis, Obando & Gironza & Panche & Aldrete, 2020

Obando, Ranulfo González, Gironza, Nancy Carrejo, Panche, Jeferson & Aldrete, Alfonso Neri García, 2020, The genus Loneura Navás (Insecta: Psocodea: ‘ Psocoptera’: Ptiloneuridae) in Colombia, description of new species and key for identification, Zootaxa 4802 (2), pp. 201-249: 217-220

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Loneura javierensis

n. sp.

Loneura javierensis   n. sp.

( Figs 40–51 View FIGURES 40–45 View FIGURES 46–51 )

Diagnosis. The only species of Loneura   , so far, with the hindwing Rs 3–4 branched. Hypandrium deeply cleft in the middle, posterior processes distally dilated, apices rounded; mesal processes curved, directed postero-laterally ( Fig. 43 View FIGURES 40–45 ). Lateral endophallic sclerites strongly curved, mesal endophallic sclerites long, robust, narrowing distally, mesal sclerite processes stout, long, slightly curved ( Fig. 45 View FIGURES 40–45 ).

Male. Color (in 80% ethanol). Body brown. Head pattern ( Fig. 42 View FIGURES 40–45 ), vertex light brown, with brown spots; front with transverse brown spots; an ochre transverse band between the inner border of each compound eye below the lower edge of the ocellar triangle to the epistomal sulcus; a cream diagonal band from the inner border of each compound eye to epistomal sulcus at the level of the antennal fossae. Compound eyes dark brown, ocelli hyaline with ochre centripetal crescents. Labrum, anteclypeus, postclypeus and mandibles dark brown. Genae dark brown. Postgenae light brown. Antennae: scape brown; pedicel and flagella pale brown, flagellomeres 1–10 with apices cream. Maxillary palps: Mx1 cream, Mx2–4 dark brown, cream basally. Tergal lobes of meso- and metathorax dark brown, with light brown edges. Thoracic pleura pale brown, with large ochre areas on mesopleura. Legs: coxae with small ochre spot basally, fore- coxae and trochanters brown; coxae, trochanters and femora of mid- and hind legs light brown; femora with two brown spots; tibiae and tarsi pale brown. Wings hyaline ( Figs 40–41 View FIGURES 40–45 ). Forewing pterostigma hyaline ( Fig. 40 View FIGURES 40–45 ), with large proximal and distal dark brown bands; brown spots on vein ends at wing margin. Hindwings ( Fig. 41 View FIGURES 40–45 ) with brown spots on vein ends at wing margin. Abdomen light brown to cream, with subcuticular ochre spots. Clunium pale brown. Hypandrium dark brown. Epiproct and paraprocts light brown.

Morphology. Head ( Fig. 42 View FIGURES 40–45 ): H/MxW: 1.39; H/d: 3.40; IO/MxW: 0.70. Vertex at the same level of the upper border of the compound eyes. Outer cusp of lacinial tips broad, with 5–6 short denticles. Mx4/Mx2: 1.29. Forewings ( Fig. 40 View FIGURES 40–45 ): L/W: 2.51. Pterostigma: lp/wp: 5.17, areola postica tall, apically rounded: al/ah: 1.36, R4+5 sinuous, M six-branched, M2, M4–6 forked or sometimes M2–6 forked and R4+5 forked. Hindwings ( Fig. 41 View FIGURES 40–45 ): l/w: 2.71; Rs three–four branched; M four–branched. Hypandrium ( Fig. 43 View FIGURES 40–45 ), posterior projections of hypandrium dilated mesally, with preapical setae on inner margin, setose as illustrated; phallosome V-shaped anteriorly, side struts slender; external parameres laminar, wide, apices rounded, bearing pores and sclerotized short teeth ( Fig. 45 View FIGURES 40–45 ); three pairs of endophallic sclerites, anterior pair thick, elongate, narrowing distally, pointed, and converging posteriorly; lateral sclerites C-shaped, wider distally; mesal pair wide and curved inwards, mesal sclerite processes distally curved outwards, elongate and truncate apically. Paraprocts ( Fig. 44 View FIGURES 40–45 ) robust, with distal setal fields and macrosetae as illustrated; sensory fields with 24–26 trichobothria on basal rosettes. Epiproct ( Fig. 44 View FIGURES 40–45 ) semioval, with short and long setae, two lateral macrosetae, a setal field on posterior border and a posterior field of microspicules.

Measurements. FW: 4675, HW: 3250, F: 1125, T: 1950, t1: 850, t2: 85, t3: 130, ctt1: 31, f1: 850, f2: 687, f3: 612, f4: 512, f5: 355, f6: 350, f7: 340, f8: 287, f9: 250, f10: 245, f11: 232, Mx4: 290, IO: 500, D: 400, d: 290, IO/d: 1.72, PO: 0.73.

Female. Color. Body, head, legs, epiproct, paraprocts and wings as in the males. Subgenital plate light brown, with pigmented area V-shaped, less pigmented posteriorly ( Fig. 49 View FIGURES 46–51 ). Gonapophyses dark brown, ninth sternum orange to yellowish.

Morphology. Head ( Fig. 48 View FIGURES 46–51 ) H/MxW: 1.42; compound eyes large, H/d: 3.64; IO/MxW: 0.72. Vertex slightly above the level of the upper border of the compound eyes; outer cusp of lacinial tips broad, with 5–6 denticles. Mx4/ Mx2: 1.32. Forewings ( Fig. 46 View FIGURES 46–51 ) L/W: 2.44; pterostigma: lp/wp: 4.49, elongate, wider in the middle; M six-branched, M2–6 forked, R4+5 forked; areola postica: la/ha: 1.58, tall, broadly triangular, apically rounded. Hindwings ( Fig. 47 View FIGURES 46–51 ) l/w: 2.72; M four-branched; Rs three-branched. Subgenital plate ( Fig. 49 View FIGURES 46–51 ) wide, setose, with posterior border rounded. Gonapophyses ( Fig. 51 View FIGURES 46–51 ): v1 elongate, slender and acuminate; v2+3 with a proximal slender heel, v2 with a row of 8–9 setae; distal process long, sinuous and acuminate, bearing microspicules on surface. Ninth sternum ( Fig. 51 View FIGURES 46–51 ) semioval, with well-defined edges anteriorly. Paraprocts ( Fig. 50 View FIGURES 46–51 ) triangular, with long and short setae; sensory fields with 30–32 trichobothria on basal rosettes. Epiproct ( Fig. 50 View FIGURES 46–51 ) triangular, wide basally, with long and short setae as illustrated.

Measurements. FW: 4757, HW: 3350, F: 1187, T: 1932, t1: 817, t2: 92, t3: 133, ctt1: 28, f1: 700, f2: 625, f3: 562, f4: 482, f5: 350, f6: 320, f7: 300, Mx4: 290, IO: 530, D: 387, d: 285, IO/d: 1.86, PO: 0.74.

Material studied. Holotype male. COLOMBIA. Magdalena. Ciénaga, San Javier , 10º52’26.9”N: 74º01’38.0”W, 1621 m. 5.IV.2016. On tree trunk. J. Mendivil & R. González. MUSENUV slide code 29313 GoogleMaps   . Paratypes: 1 female, same data as the holotype. MUSENUV slide code 29314 GoogleMaps   . 1 male, Magdalena, National Natural Park Tayrona , 11º20’N: 74º2’W, 700 m, IAvH E-163777, 19.VII.2002. Malaise trap GoogleMaps   .

Etymology. The specific epithet refers to San Javier, Magdalena Department where the types were found.


Departamento de Geologia, Universidad de Chile