Fannia fusconotata ( Rondani, 1868 )

Dominguez, Martha Cecilia, 2007, A taxonomic revision of the Southern South American species of the genus FANNIA Robineau-Desvoidy (Diptera: Fanniidae), Papéis Avulsos de Zoologia (São Paulo) 47 (24), pp. 289-347 : 312-315

Fannia fusconotata ( Rondani, 1868)

Figs. 1D , 8 A-G

Myiantha fusconotata Rondani, 1868:27 .

Fannia fusconotata ; Stein, 1919:131, 167; Mazza & Joerg, 1939:43; Mazza & Oribe, 1939:66; James, 1947:130; Pont, 1972:4; Carvalho et al., 1993:10; 2003:12.

Diagnosis: Small yellow and grey species. Frontal vitta at narrowest point reduced to a thin line. Eye bare. Parafacial bare. Antenna light brown and orange-yellow. Palpus orange-yellow. Thorax grey-brown, thin brown vittae visible in full length of acr and at base of dc lines. Two strong pra near spal. T 2 constricted in basal half, lacking preapical protuberance. F3 normal with a preapical posteroventral protuberance only evident in posterior view; with 1 row of very short setulose pv and a subapical tuft of short hair-like pv. T 3 lacking medial row of ad. Wing clear. Halter yellowish-white. Abdomen heart-shaped, grey-brown in ground colour, heavily yellow pruinose on laterals of tergites 2-4.

Type material examined: Holotype n: Argentina, Mendoza. Rondani ( MZLS).

Other material examined: Argentina. Mendoza. 44 o, 31 n, Ñacuñan, I / III-1997, col Guillermo Debandi ( IADIZA). 2 o, Lavalle, Telteca, XII-1997 / II-1998, col. Sergio Roig. ( IADIZA). 17 n, Uspallata, Arroyo San Alberto, 29-XII-2000, col. Domínguez ( IADIZA). 2 o, 21 n, Uspallata, desembocadura Arroyo Uspallata, XII-2000 / II-2001, col. Domínguez ( IADIZA). 27 o, 10 n, Lavalle, Costa de Araujo, XI-2001, col. Domínguez ( IADIZA). 4 n, Potrerillos, Quebrada de los Gateados, XII-2001, col. Domínguez ( IADIZA). 4 n, Uspallata, Quebrada del Río Seco , XII-2001, col. Domínguez ( IADIZA). 15 n , Mendoza, Río Mendoza

Km 1137, XII-2001, col. Domínguez ( IADIZA). 8 n, Mendoza, Ciudad. CRICYT, X-2001, col. Domínguez ( IADIZA) .

Description

Male: body length 3.2-3.5 mm

Head ( Fig. 1F ): Frontal vitta black, at narrowest point reduced to a thin line. Fronto-orbital plate heavily silvery grey pruinose, at uppermost pair of fr 1.5x width of anterior ocellus. Six to seven fr, slightly shorter than flagellum. Eye bare, antero-internal facets 2x diameter of remaining; pocl of uniform length. Face and parafacial silvery grey; parafacial at base of flagellum 0.5x width of same and bare. Facial carina dark grey. Gena grey. Scape light brown, pedicel and base of flagellum orange-yellow; flagellum short and broad 1.2 to 1.7x as long as broad, heavily silvery grey pruinose. Arista light brown at base, black at apex and short pubescent. Palpus orange-yellow, slightly spatulated, the apex 2x width of base. Lower oral margin, slightly concave and grooveless.

Thorax: Scutum grey-brown, thin brown vittae visible in full length of acr and at base of dc lines, also visible at base of scutellum; pleura grey; 3+3 acr, 1 prescutellar acrs; 2+3 dc; pprn with few setulae; 2 strong pra near spal; prealar area bare; 2 prepm, lacking surrounding setulae.

Legs: Coxae brown, trochanters yellow, femora brown; tibiae yellow at apex and base, remaining brown, tarsi dark-brown to black. F1 normal (length/width = 4.7); 1 row of pd as long as femoral width; 1 row of p as long as femoral width; 1 row of pv short in apical fourth but 8 to 9 longer pv at apex. T 1 with 1 subapical pd; 1 apical v and pv; 1 submedial and 1 apical ad. Fore tarsomere 1 0.3x length of fore tibia, tarsomeres 4 and 5 as wide as long. F2 with 1 row of ad; 2 rows of a, one in full length and other in medial 1/3; with 1 row of av sparse at base, denser and shorter at apex; 2 rows of pv, upper row as long as femoral width and lower short at base, longer at apex; 1 row of p longer and ventrally directed at apex. T 2 constricted in basal half, lacking preapical protuberance. Ventral pubescence sparse, as long as femoral width; 1 submedial and 4 apical a; 1 subapical ad; 1 strong apical av-v; 1 submedial and 1 apical pd; 1 apical pv and p. C3 bare at apex of posterior margin. F3 normal (length/width = 5.4), with a preapical posteroventral protuberance only evident in posterior view; 1 row of ad longer and ventrally directed towards apex; 1 row of very short av, and 2 preapical av longer than femoral width; with a row of very short setulose pv and a subapical tuft of short hair-like pv, slightly longer than femoral width ( Fig. 11A, B ). T 3 with 1 submedial and 1 subapical d; 1 submedial and 1 apical ad, lacking medial row; 2 to 3 medial and 1 apical short av; apex of ventral and posteroventral surface with a weak ctenidium.

Wing: Clear. Lower calypter oval, upper rounded, both white. Knob of halter yellowish-white, light brown at base.

Abdomen: Heart-shaped, tergite 2 over 2x width of tergite 4. Grey-brown in ground colour, heavily yellow pruinose on laterals of tergites 2-4. Hypopygium not protruding.

Postabdomen: Posterior margin of sternite 5 profoundly indented and covered with setae ( Fig. 11C ). Hypandrial arms directed outwards and expanded at apex; postgonites inconspicuous; aedeagus membranous ( Fig. 11D ). Epandrium slightly broadened at posterior margin; cercal plate fused, slightly bilobated at base; bacilliform process cork-screw shaped; surstily long and simple, marginal respect to epandrium ( Fig. 11E, F ).

Female: body length 3.0- 3.4 mm. Differs from male as follows:

Head: Frontal vitta brown. Ocellar triangle extending up to lowemost fr. Frons and vitta broad, distance between eye margins wider than 0.33x width of head. Four to five strong fr, shorter than flagellum, with shorter intersticial setulae. Upper and lower orb present, with a single row of fronto-orbital setulae; pocl less than 0.5x length of outer and inner verticals; the latter longer than in male.

Thorax: With two rows of presutural acr, and a third row before transverse suture.

Legs: F2 with 1 row of very short ad; posteroventral row absent. T 2 straight, cone-shaped, the base narrower than apex; ventral pubescence absent; with 1 very long apical and 1 submedial av-v; 1 apical p; 1 apical and 1 subapical pv. C3 bare at apex of posterior margin. F3 lacking preapical protuberance on ventral surface; posterior and posteroventral surface bare. T 3 with 1 submedial ad setae, lacking apical setae.

Abdomen: Heart-shaped, tergite 2 more than 2x width of tergite 5. Grey in ground colour, light brown-yellowish at apex of tergites 1 and 2; tergites 2-5, trimaculated, with lateral and central longitudinal brown markings in posterior half.

Postabdomen: Cerci normal, short; anal plate as wide as broad and uniformly covered with setae; sternite 8 reduced to two anterior plates bearing 2 setae and 4 to 5 setulae; postabdominal spiracles 7 and 8 on tergite 6 ( Fig. 11G ). Two wrinkled pear shaped spermathecae, with partially sclerotized ducts ( Fig. 11H ).

Distribution: NEOTROPICAL: province of Mendoza, Argentina.

Figures

FIGURE 1: F. coxata Shannon & Del ponte. A. Male hind coxa and abdomen, lateral. F. schnusei Stein. B. Male fore tarsi, dorsal. C. Male hind leg, anterior. F. fusconotata (Rondani). D. Male head, anterior. F. scalaris (Fabricius). E. Male mid coxa, anterior. F. roigi sp. nov. F. Male mid tibia, posterior. G. Male hind leg, anterior. F. hermani sp nov. H. Wing.

FIGURE 8: Fannia fusconotata (Rondani). A. Male hind leg, anterior. B. Male hind leg, posterior. C. Male sternite 5, ventral. D. Male hypandrium, ventral. E. Male terminalia external structures, ventral. F. Female terminalia external structures, ventral. G. Spermathecae.

FIGURE. 11: Fannia hirtifemur (Stein). A. Male hind leg, anterior. B. Male sternite 5, ventral. C. Male hypandrium, ventral. D. Male terminalia external structures, ventral. E. Male terminalia external structures, lateral. F. Spermathecae.