Chrysoclista germanica, Šumpich & Huemer, 2016

Šumpich, Jan & Huemer, Peter, 2016, Chrysoclista germanicasp. nov. and C. gabretica stat. nov., with an updated checklist of the genus (Lepidoptera: Elachistidae: Parametriotinae), Acta Entomologica Musei Nationalis Pragae 56 (2), pp. 845-854 : 850-852

Chrysoclista germanica sp. nov.

( Figs 4 , 8 , 16 )

Type locality. Germany, Thuringia, Bad Blankenburg.

Type material. HOLOTYPE: J, ‘ 4.VII.1977 c [handwritten] / Blankenburg / Buntsandstein / Dr. Steuer’ ( ZMHB).

Diagnosis. Externally similar to C. linneella , C. abchasica and C. gabretica but slightly darker with reduced orange pattern and broader blackish-brown edge of the fore wings, particularly in the dorsal and terminal part, with both rounded dots integrated ( Fig. 4 ). Significant differences are found in the labial palpi which are comparatively short, only slightly curved and predominantly silvery. In the male genitalia the shape of the gnathos is similar to C. linneella ( Fig. 17 ), whereas the shape of the anellus lobes resembles C. splendida Karsholt, 1997 and the Nearctic C. villella (Busck, 1904) , though the combination of these characters is unique for C. germanica sp. nov. Moreover, the shape of the valva is unmistakable.

Description. Adult. Head and face covered by dark bronze scales with metallic shine. Labial palpi comparatively short, only slightly curved, covered with silvery scales, the third segment predominantly brown on ventral surface. Antennae filiform, brown with yellow tip. Wingspan 12 mm. Ground colour of fore wings dark orange, very broad dark brown edge particularly in the apical and terminal part. In the orange area three rounded silver-dusted black spots are present, one in the middle of the costal margin and two in the dorsum, together creating an almost regular triangle. Fringes dark brown in the colour of the edge. Hindwings dark brown, fringes long, in colour of wings.

Male genitalia. Vinculum broad, rounded. Uncus comparatively narrower. Dorsal processes in distal part of uncus small, rounded. Gnathos arms nearly regularly shaped, short and broad with approximately two dozens of thorns distally. Valva comparatively narrow, distinctly curved ventro-proximally, rounded terminally. Phallus pronouncedly curved, narrow, with a cluster of approximately two dozens of comparatively short spines in its terminal part.Anellus lobes elongated, digitate, rounded with numerous small bumps apically.

Female genitalia. Unknown.

Etymology. The species epithet is Latin adjective germanicus (- a, - um), given after the country of origin of the holotype ( Germany).

Biology. Unknown.

Distribution. Germany (Bad Blankenburg) (this paper).

C. abchasica ( Sinev, 1986) Georgia (Abkhazia)

C. ankaraensis Seven, 2014 Turkey (Ankara)

C. cambiella (Busck, 1915) U.S.A. (Oregon, Idaho, Montana), Canada (British Columbia, Alberta) C. gabretica Šumpich, 2012 stat. nov. Austria, Czech Republic, Italy

C. germanica Šumpich & Huemer , sp. nov. Germany

C. grandis Koster, 2002 U.S.A. (California, Colorado)

C. lathamella Fletcher, 1936 Central and Nothern Europe

= C. bimaculella sensu HAWORTH (1828) ,

nec THUNBERG (1794)

= C. razowskii Riedl, 1965

C. linneella (Clerck, 1759) Europe, U.S.A. (eastern part), Canada

= C. schaefferella Duponchel, 1828

= C. gemmatella Costa, 1836

C. splendida Karsholt, 1997 Central Europe

= C. bimaculella auct.

= C. razowskii auct.

C. villella (Busck, 1904) U.S.A. (Washington), Canada (British Columbia)

C. zagulajevi Sinev, 1979 Georgia (Adzharia)

Fig. 18. Neighbour-joining tree (Kimura 2 parameter, built with MEGA 6; cf. TAMURA et al. 2013). Source: DNA Barcode data from BOLD (Barcode of Life Database, cf. RATNASINGHAM & HEBERT 2007).

C. basiflavella Matsumura, 1931 ( Japan) junior synonym of Stathmopoda auriferella (Walker, 1864) in Stathmopodidae ( MORIUTI 1975)

C. bicolorella Matsumura, 1931 ( Japan) unplaced species in Tineidae : Hieroxestinae (Sinev, pers. comm.)

C. hexachrysa Meyrick, 1935 ( Japan) Pancalia hexachrysa (Meyrick, 1935) in Cosmopterigidae (KU- ROKO 1982)

C. hygrophilella Viette, 1957 ( Réunion Island) Stathmopoda hygrophilella (Viette, 1957) in Stathmopodidae ( SINEV 2015)

C. monotyla Meyrick, 1921 ( Australia) unplaced species in Chrysopeleiidae (Sinev, pers. comm.)

C. thrypsiphila Meyrick, 1912 ( Ceylon) unplaced species in Chrysopeleiidae (Sinev, pers. comm.)

C. trilychna Meyrick, 1928 ( India) unplaced species in Chrysopeleiidae (Sinev, pers. comm.)

Figures

Figs 1–5. Voucher specimens of Chrysoclista, males. 1–3 – C. gabretica Šumpich, 2012: 1 – Czech Republic, holotypus (10.5 mm); 2 – Italy (12 mm); 3 – Austria (12 mm). 4 – C. germanica sp. nov., holotypus (12 mm). 5 – C. linneella (Clerck, 1759), Poland, Wrocław, 1.viii.1881, ex. coll. O. Staudinger (ZMHB) (12.5 mm).

Figs 6–10. Heads of Chrysoclista species.6–7 – C. gabretica Šumpich, 2012:6 – Italy; 7 –Austria.8 – C. germanica sp. nov. 9–10 – C. linneella (Clerck, 1759): 9 – Czech Republic, Prag (= Praha), 1870, ex. coll. F. A. & O. Nickerl (NMPC); 10 – Prag, 22.viii.1940, V. Vlach leg. (NMPC).

Figs 16–17. Genitalia of Chrysoclista species. 16 – C. germanica sp. nov., holotypus, general view (a–b – two different views of the terminal part of phallus with apical thorns, c – anellus lobe, d – gnathos). 17 – C. linneella (Clerck, 1759), detail of gnathos, England, ex. coll. Bang-Haas (ZMHB).