Agyrtodes globosus Seago, 2009

Seago, Ainsley E., 2009, Revision Of Agyrtodes Portevin (Coleoptera: Leiodidae), The Coleopterists Bulletin (mo 7) 63, pp. 1-73 : 37-38

publication ID	https://doi.org/ 10.1649/0010-065x-63.sp7.1
publication LSID	lsid:zoobank.org:pub:B58B2216-0CFA-41C6-8141-7024E65ECF85
DOI	https://doi.org/10.5281/zenodo.4912299
persistent identifier	https://treatment.plazi.org/id/039887BA-FFC3-C068-92E7-80B6EEC7917A
treatment provided by	Carolina
scientific name	Agyrtodes globosus Seago
status	sp. nov.

Treatment

Agyrtodes globosus Seago View in CoL , new species

(Figs. 10, 47, 67, 87, 101)

Holotype: male, labeled ‘‘ AUSTRALIA: n. Qld , Tully Falls S. F. 1,000 m, 9. 5 km SSW Ravenshoe, 7. XII. 87– 7. I. 1988, Storey & Dickinson / MDPI Intercept Trap, Site No. 29A’’ (ex QDPI, deposited in QMBA).

Paratypes: listed in Appendix 1 (100 specimens)

Distribution. Australia: Northeast Queensland, Atherton Tablelands ( Fig. 101 View Fig ).

Diagnosis. This species can be recognized by its large, round, convex body form, uniform brown to light brown coloration, and the lack of impressed longitudinal striae on the elytra. Agyrtodes crassus and A. eucalypti (both known only from New South Wales) have a similarly large and ovoid body shape, but can be distinguished from A. globosus by a red-and-black dorsal color pattern and swollen apical maxillary palpomeres ( A. crassus ) or by a narrower, posteriorly tapered body shape, darker brown pronotum, and the presence of longitudinally impressed elytral striae ( A. eucalypti ). The short, triangular median lobe and long, flattened, oar-shaped parameres of A. globosus also distinguish it from A. crassus and A. eucalypti (both of which have an elongate, slender median lobe).

Description. TBL 5 2.2 mm, EW 5 1.4 mm, PNW 5 1.3 mm, HW 5 0.65 mm. (TBL variation: 2.1–2.4 mm).

Body shape (Fig. 10) large, broad, wider than other Agyrtodes with the exception of A. crassus . Elytra strongly convex, nearly hemispherical. Color uniformly dark reddish brown, fading to medium brown in older or teneral specimens. Dorsum evenly pubescent, vestiture composed of long, fine, golden hairs.

Head broad, dark reddish-brown; eyes very large and bulging, epistomal suture absent. Frons and clypeus with loose vestiture of fine, stiff golden setae, integument shining, setal insertions in small, fine punctures. Clypeus short, transverse; labrum longer than clypeus, equally wide, rectangular in shape with anterior margin very weakly excavate. Maxilla with slender galea with weak apical brush, wide lacinia with broad, well-sclerotized spore-brush with small spines arranged in parallel rows; maxillary palp with apical segment lanceolate, weakly attenuate at apex, 1.5 3 as long and slightly broader than penultimate segment. Labium with four long digitiform sensillae arranged medially on anterior margin of ligula, labial palpi three-segmented, apical segments with basal clusters of minute digitiform sensillae. Antennae ( Fig. 47 View Figs ) with basal segments slender, club robust; segments 1–3 light brown, 4–6 medium brown, 7–11 dark brown. Segments 1 and 2 broad, 3–6 slender, cylindrical, gradually decreasing in length; segment 6 very slightly wider and approximately twothirds as long as segment 3. Segment 7 elongate, approximately 2 3 length of 8, narrower than 9–11; segment 8 subtly elongate, ovoid, slightly wider at apex. Segments 9–11 longer than wide, apically expanded, broader than all preceding segments. Segment 12 ovoid, with weakly sclerotized, gently attenuate apex.

Pronotum shining, brightly reflective, integument extremely smooth with no trace of microsculpture; minute, shallow punctures present only at setal insertions. Hind angles square, very feebly produced posteriorly; entire hind margin of pronotum transparent.

Elytra convex, uniformly reddish-brown, with conspicuous, deeply punctate transverse strigae; no trace of longitudinal strial impressions. Integument shining, vestiture primarily composed of semirecumbent fine, dark golden setae, but also with 4–5 sparse longitudinal rows of light golden erect setae. Elytral apices rounded, very bluntly pointed. Sutural striae shallow, basally effaced. Scutellum light reddish-brown.

Pronotum short, weakly convex, procoxal cavities large and globular, trochantins broadly exposed. Mesosternal carina strongly elevated, highest anterior to mesocoxal separation; mesepimera elongate, bluntly rounded at mesocoxal articulation. Metepisterna concave, elongate, bluntly triangular, broad anteriorly and narrowing posteriorly. Metasternum flat to concave, not convex, with fine, confused striolation and shallow setiferous punctures.

Legs robust, of typical proportions; femora slightly flattened. Meso- and metatibia in both sexes armed with two external rows of unusually large, dark spines; protibial armature normal. Male mesotibia gently curved in distal third, with mesotibial spines halting abruptly before bend. Male protarsi with segments 1–3 expanded and bearing tenent setae; male mesotarsi with first segment bearing small, dense cluster of tenent setae, mesotarsi not expanded. Claws and female tarsi simple.

Male genital segment ( Fig. 67 View Figs ) broadly conical, sternum X well-sclerotized and fused ventromedially to pleurites; anterior apophysis slender and produced posteriad, rounded apically. Aedeagus ( Fig. 87 View Figs ) with median lobe extremely short, penis triangular, broad at base, tapering smoothly to bluntly pointed apex. Basal piece short, broad. Parameres extending far past apex of median lobe; flattened, blade-like with slight expansion halfway along length and at apices. Interior faces of parameres with abundant setiferous punctures; apices with small patches of stout, conical teeth; each apex with one terminal and one slightly subapical long seta. Endophallus with abundant small spines and 5+ large, curved, fang-shaped sclerites. Female terminalia with coxites long, basally connate; styli less than 1/3 as long as coxites, slender, each with long terminal seta.2 3 as long as stylus.

Natural History. Frequently collected in series in high-altitude (1,000+ m) flight-intercept traps and pyrethrum fogging of logs and tree bases in wet forests.

Etymology. The specific epithet globosus is a regular Latin adjective, used here in reference to the large, rounded body shape that distinguishes this species.