Mycomya klossi Edwards, 1931

Väisänen, Rauno, 2014, New Mycomya species from South-East Asia (Diptera, Mycetophilidae), Zootaxa 3815 (4), pp. 526-540 : 529

Mycomya klossi Edwards, 1931

Figs. 2 A–C

Mycomya klossi Edwards, 1931 . Edwards 1931a: 488

Material studied. Holotype. ♂. MALAYSIA, N. Borneo, Bettotan. nr. Sandakan Aug. 18th 1927. 2458. (NHM).

Description. ♂. Head. Palp, other mouthparts, face and posterior parts of head yellow, ocellar prominence brown. Antenna yellowish, scape, pedicel and base of 1st flagellomere yellow. 1st flagellomere about 1.5x its width, 2nd flagellomere 1x its width. Thorax. Pronotum yellow, with 4 long setae. Scutum yellow to yellowish without distinct stripes. Anepisternum and preepisternum yellow. Scutellum brownish, with 4 long setae. Laterotergite yellowish. Mediotergite yellowish, bare. Wing. Length 2.5 mm. Wing hyaline, small cell and wing apex weakly infuscated. Sc ending in R1 distad of middle of small cell, Sc1 missing. Apical part of Sc bearing 9 small setae. Small cell about 2x as long as wide. Cu fork slightly proximal to M fork. M ratios: 0.80, 1.00. Cu ratios: 0.67, 0.93. Small setae: M petiole: 0; M1: 0; M2: 0; Cu petiole: 0; Cu1: 0; Cu2: 0. Halter yellowish, apex brown. Legs. Coxae and femora yellow, tibiae and tarsi brownish. Coxa 2 without spur. Leg ratios: bt1:t1 = 0.82, bt2:t2 =?, bt3:t3 = 0.55. Abdomen. Tergites 1–5 yellow, with brownish posterior margins, 6–7 brownish. Sternites 1–5 yellow, 6–7 brownish. Hypopygium. Figs. 2 A–C, yellow. Sternal lateral appendage relatively long, apically rounded, its subapical part about as wide as basal part, densely setose ( Fig. 2 C). More than apical ½ of sternal synsclerite covered by long setae. Gonostylus with two teeth. Sternal submedian filament not visible due to poor condition of hypopygium.

Discussion. Edwards (1931) noted that in M. klossi “the venation and other characters indicate an affinity with the group of M. fimbriata Mg. ”, and based on his comment, Väisänen (1984a) tentatively placed M. klossi in the subgenus Neomycomya Väisänen (the type species of which is Sciophila fimbriata Meigen , i.e. M. fimbriata ). However, M. klossi clearly belongs to the widely distributed subgenus Cymomya , based on the presence of the medial comb in the tergal part of the hypopygium lacking the tergal lateral appendages, and of the sternal synsclerite divided into two large lateral parts bearing wide, setose lateral appendages ( Väisänen 1984a). Mycomya klossi is smaller (wing length 2.5 mm) than the other Oriental species of the subgenus, very pale, and with relatively conspicuous wing markings compared to the other Mycomya species, resembling those in the genus Neoempheria . The long, apically rounded shape of the sternal lateral appendage, with dense setosity, is diagnostic to M. klossi ( Fig. 2 C). It is closely related to M. paraklossi sp. n., another yellowish species, with only the posterior margins of abdominal tergites 1–5 brown and large setose sternal lateral appendages. However, M. paraklossi is bigger than M. klossi and the subapical part of its sternal lateral appendage is much wider that the basal part ( Fig. 5 C). Mycomya aonyx Väisänen from Myanmar is also quite similar to M. klossi , but its sternal lateral appendage is D-shaped ( Väisänen 2013a: fig. 4B) and not as symmetrically rounded as in M. klossi , it lacks the conspicuous wing markings, and it has a darker coloration, with abdominal tergites 1–5 brownish to brown, sometimes with yellowish anterolateral spots. The holotype of M. klossi was collected in a lowland area in Sabah, northern Borneo, in August by Messrs. C. Boden Kloss and H. M. Pendlebury. During their visit to British North Borneo in July–September, 1927 no ascents were made above 200 ft. (Edwards 1931). A large proportion of the biologically very diverse lowland forest in the Sandakan region has been recently converted into oil palm plantations.

Figures

FIGURE 2. Mycomya klossi Edwards, apical part of male hypopygium. A – B tergal and sternal views, C sternal lateral appendage. Scale 0.2 mm.

FIGURE 5. Mycomya paraklossi sp. n., male hypopygium. A – C tergal, sternal and lateral views, D apex of aedeagus, twice enlarged, E tergite 8. Scale 0.2 mm.