Stegana (Steganina) ornatipes

Cheng, Yu, Gao, Jian-Jun & Chen, Hong-Wei, 2009, Stegana ornatipes species group from the Oriental Region (Diptera: Drosophilidae), Zootaxa 2216 (1), pp. 37-48 : 38

Stegana (Steganina) ornatipes species group

Diagnosis. Surstylus large, with 1 strong prensiseta apically and several thin, long setae ( Figs 2 , 8 , 12 , 16 , 20 , 24 , 28); 10th sternite mostly narrowed, nearly arcuate, with 1 pair of projections posterolaterally ( Figs 3 , 9 , 13 , 17 , 21 , 25 , 29); gonopods (dorsal arch in Bächli et al. 2004) with 1 pair of projections sublaterally ( Figs 4 , 10 , 14 , 18 , 22 , 26 , 30); aedeagus mostly separated from basally and with cilia ( Figs 4 , 10 , 14 , 18 , 22 , 26 , 30).

Description. Male and female: Frons and face not rectangular in profile. Eyes red. Ocellar triangle black, with 1 pair of small setae above ocellar setae. Postvertical setae slightly behind vertex ridge. Frons shiny, brown, with sporadic, minute setulae submedially, and a black, transverse band above ptilinal fissure. Proclinate orbital setae slightly nearer to ptilinal fissure than to inner vertical setae. Face black, with yellow, transverse band medially, broadened ventrally; facial carina absent. Clypeus black medially, yellow laterally. Palpus with 1–2 longer setae distally and several shorter setae basally. Gena yellow, narrow (ch/o <0.10). Vibrissa prominent; other orals small. Occiput glossy, yellow, but black around occipital foramen. Mesonotum brown to dark brown, mostly unicolorous. Mesopleuron with a black longitudinal stripe above (running from propleuron to base of halter). Postpronotal lobe brown on upper part, white on lower part, with 1 long and a few short setae. Acrostichal setulae approximately in 10 irregular rows. Prescutellar setae 1 pair. Scutellum brown to dark brown; basal setae divergent; apical setae crossing with each other. Wing dark brown anteriorly, pale posteriorly, curved downward on distal part. Basal medial-cubital crossvein present. C 1 with 2 isometric setae. Costal vein with 6–9 minute spinules on ventral surface between R 2+3 and R 4+5. R 2+3 obviously curved to costa at tip; R 4+5 and M 1 convergent distally. Halters white basally, grey-brown distally. Fore femur with 1 row of setae on posterior surface and 1 row of 3–6 setae on ventral surface. Apical seta present on mid tibia. Preapical dorsal setae present on all tibiae. Mid tarsus with 4–6 strong setae on basal part of dorsal surface. Mid and hind tarsomeres with 2 and 1 row(s) of minute cuneiform setulae on ventral surface, respectively; fore and hind 1st tarsomeres slightly shorter than the rest combined; mid 1st tarsomere longer than the rest combined. All abdominal tergites dark brown to black. Sternites brown; 3rd to 5th broadened; 6th covered with 5th.

Male terminalia: Epandrium with setae and pubescence ( Figs 1 , 7 , 11 , 15 , 19 , 23 , 27). Surstylus separated from epandrium, with thin, long setae on margins of outer surface and inner surface ( Figs 2 , 8 , 12 , 16 , 20 , 24 , 28). Cercus separated from epandrium, setigerous, sometimes with sporadic pubescence ( Figs 1 , 7 , 11 , 15 , 19 , 23 , 27). Hypandrium anteromedially protruded, connected with base of aedeagus by articulating plate like paramere ( Sidorenko 1997, 1998) but mostly lacking sensilla ( Figs 4 , 10 , 14 , 18 , 22 , 26 , 30). Paramere (paraphysis in Bächli et al. 2004) absent ( Figs 4 , 10 , 14 , 18 , 22 , 26 , 30). Gonopods forming posteromedian lobe, sclerotized, laterally contiguous to posterior ends of hypandrium ( Figs 4 , 10 , 14 , 18 , 22 , 26 , 30). Aedeagal apodeme slender, contiguous to base of aedeagus ( Figs 4 , 10 , 14 , 18 , 22 , 26 , 30).

Female terminalia ( Figs 5, 6 ): Epiproct and cerci not pubescent; oviscapt (8th sternite) with numerous, minute spinules; hypoproct with numerous setae and pubescence.

Figures

FIGURES 1–6. Stegana (Steganina) albiventralis sp. nov., ♀: 1–4 male terminalia, 5, 6 female terminalia. 1. Epandrium (epand), surstylus (sur) and cercus (cerc) (lateral view); 2. surstylus (frontal view); 3. 10th sternite (ventral view); 4. hypandrium (hypd), gonopods (gon), aedeagus and aedeagal apodeme (ventral view); 5. epiproct (epi) and cercus (cerc) (dorsal view); 6. 8th sternite (ovi) and hypoproct (hypo) (Scale bars: 0.1 mm).

FIGURES 7–10. Stegana (Steganina) angustigena sp. nov., ♂: 7. Epandrium, surstylus and cercus (lateral view); 8. surstylus (frontal view); 9. 10th sternite; 10. hypandrium, gonopods, aedeagus and aedeagal apodeme (ventral view) (For abbreviations see Figures 1–6. Scale bars: 0.1 mm).

FIGURES 11–14. Stegana (Steganina) lingnanensis sp. nov., ♂: 11. Epandrium, surstylus and cercus (lateral view); 12. surstylus (frontal view); 13. 10th sternite; 14. hypandrium, gonopods, aedeagus and aedeagal apodeme (ventral view) (For abbreviations see Figures 1–6. Scale bars: 0.1 mm).

FIGURES 15–18. Stegana (Steganina) mengla sp. nov., ♂: 15. Epandrium, surstylus and cercus (lateral view); 16. surstylus (frontal view); 17. 10th sternite; 18. hypandrium, gonopods, aedeagus and aedeagal apodeme (ventral view) (For abbreviations see Figures 1–6. Scale bars: 0.1 mm).

FIGURES 19–22. Stegana (Steganina) nulliseta sp. nov., ♂: 19. Epandrium, surstylus and cercus (lateral view); 20. surstylus (frontal view); 21. 10th sternite; 22. hypandrium, gonopods, aedeagus and aedeagal apodeme (ventral view) (For abbreviations see Figures 1–6. Scale bars: 0.1 mm).

FIGURES 23–26. Stegana (Steganina) pilosella sp. nov., ♂: 23. Epandrium, surstylus and cercus (lateral view); 24. surstylus (frontal view); 25. 10th sternite; 26. hypandrium, gonopods, aedeagus and aedeagal apodeme (ventral view) (For abbreviations see Figures 1–6. Scale bars: 0.1 mm).