Mortoniella redunca, Blahnik & Holzenthal, 2008

Blahnik, Roger J. & Holzenthal, Ralph W., 2008, Revision of the Mexican and Central American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae), Zootaxa 1711 (1), pp. 1-72 : 49-52

Mortoniella redunca , new species

Fig. 25

As previously discussed, Mortoniella redunca is similar to a group of species, including M. carinula , M. opinionis , M. papillata , M. sicula , n. sp, and M. umbonata , n. sp., all distinguished by a combination of characters, including inferior appendage invaginated mesally and with protruding, often apically acute, apicolateral projections, and also a phallicata with well developed dorsolateral processes. Among these species, M. redunca is recognized by the unique structure of its dorsal phallic spine, which has the apex formed into a characteristic downward hook. The paramere appendages of this species are somewhat variable in length, but tend to be shorter than in most of the other species in this group; the apices are scabrous. Other useful characters to note are that the endophallic spine is less curved than in most members of this group and the apicomesal process of tergum X is scarcely developed.

Adult. Length of forewing: male 3.2–3.8 mm, female 3.8–4.7 mm. Forewing with forks I, II, and III, hind wing with forks II and III. Overall color medium brown, palps darker brown, apices of tibiae, tarsal and basal antennal segments light brown. Wing bar at anastamosis indistinctly marked with pale, light brown setae. Ventral process of abdominal segment VI (male) short, ventrally oriented, subtriangular, subacute apically, not constricted basally.

Male genitalia. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin forming rounded projection in dorsal half, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, dorsal excision narrow, much less than half width of segment. Tergum X without, or with very weakly developed, mesal projection and projecting lateral lobes, lateral lobes elongate, narrow as viewed dorsally, sclerotized and subacute apically, slightly mesally curved. Inferior appendage without apicomesal projection, apicolateral projections elongate, tapering, acuminate, apices slightly dorsally curved; appendage nearly linear basolaterally, only slightly projecting on each side; mesal pockets of inferior appendage with apical processes short, dorsally curved. Paramere appendages nearly uniform in width, dorsally curved from base, posteriorly directed apically, apices with scabrous development, apex acute. Dorsal phallic spine distinctive in shape, dorsal margin sinuous, apex, in lateral view, distinctly widened and hooked downward; as viewed dorsally, with apex narrowed, nearly acute, sometimes slightly asymmetrically curved. Phallicata relatively short, ventral margin distinctly curved and produced, dorsolaterally with prominent, paired, processes on either side, each more or less rounded and laterally explanate. Endophallic membrane with single prominent spine, sometimes sharply curved at base, apex nearly straight.

Holotype male: COSTA RICA: Alajuela: Río Toro, 3.0 km (road) SW Bajos del Toro, 10°12'14"N,

084°18'58"W, 1530 m, 3–4.ix.1990, Holzenthal, Blahnik & Huisman ( UMSP000001261 ) (pinned) ( UMSP) .

Paratypes: COSTA RICA: Alajuela: same locality and date as holotype — 11 males, 20 females (pinned), 10 males (alcohol) ( UMSP) ; same locality, 11.ii.1992, Holzenthal, Muñoz & Kjer — 1 male, 3 females (pinned) ( INBIO) , 4 males, 2 females (alcohol) ( UMSP) ; Puntarenas: Río Cotón, in Las Alturas , 08°56'17"N, 082°49'34"W, 1360 m, 16.ii.1986, Holzenthal, Morse & Fasth — 3 males (alcohol) ( UMSP) ; Río Bellavista , ca. 1.5 km NW Las Alturas, 08°57'04"N, 082°50'46"W, 1400 m, 18.ii.1986, Holzenthal, Morse & Fasth — 2 males (pinned) ( NMNH) ; same locality, 8–9.iv.1987, Holzenthal, Hamilton & Heyn — 1 male (alcohol) ( UMSP) ; same locality, 2–3.viii.1987, Holzenthal, Morse & Clausen — 3 males, 1 female (pinned), 1 male (alcohol) ( UMSP) ; same locality, 10–11.viii.1990, Holzenthal, Blahnik & Muñoz — 2 males, 51 females (pinned), 2 males (alcohol) ( UMSP) ; same locality, 16–17.iii.1991, Holzenthal, Muñoz & Huisman — 3 males, 2 females (pinned), 37 males (alcohol) ( UMSP) ; Zona Protectora Las Tablas, Río Cotón , Sitio Cotón, 08°56'28"N, 082°47'13"W, 1460 m, 15.iv.1989, Holzenthal & Blahnik — 1 male (alcohol) ( UMSP) ; San José: Río Chirripó Pacífico , 9.5 km NE Rivas, 09°28'12"N, 083°35'28"W, 1370 m, 23.ii.1986, Holzenthal, Morse & Fasth — 1male, 4 females (pinned), 12 males (alcohol) ( UMSP) .

Etymology. This species is named M. redunca , from the Latin word reduncus, meaning curved back, and referring to the recurved apex of the dorsal phallic spine of this species.

Figures

FIGURE 25. Mortoniella redunca, new species. Male genitalia: A—lateral; B—segment IX and tergum X, dorsal; C— phallic ensemble, ventral; D—dorsal phallic spine, dorsal.