Khamisia banisad Saaristo and van Harten

Platnick, Norman I. & Berniker, Lily, 2015, The Goblin Spider Genus Khamisia and Its Relatives (Araneae, Oonopidae), American Museum Novitates 2015 (3837), pp. 1-68 : 8-21

publication ID

https://doi.org/ 10.1206/3837.1

persistent identifier

https://treatment.plazi.org/id/039A879C-A616-6F25-FE76-4E00FD29FB81

treatment provided by

Carolina

scientific name

Khamisia banisad Saaristo and van Harten
status

 

Khamisia banisad Saaristo and van Harten View in CoL

Figures 97–100 View FIGURES 90–100. 90–96

Khamisia banisad Saaristo and van Harten, 2006: 136 View in CoL , figs. 18–21 (female holotype from Khamis Bani Sa‘ad, Yemen, in MZT; examined).

DIAGNOSIS: Females resemble those of K. hayer but have a longer median genitalic process (fig. 100).

MALE: Unknown.

FEMALE (PBI_OON 9870, figs. 97–100): Total length 1.53. Anterior genitalic process long, distally widened, lateral processes with recurved, rounded tips that may serve as spermathecae.

MATERIAL EXAMINED: YEMEN: Al Mahwit: Khamis Bani Sa‘ad , June 23, 1999, leaf litter in and around banana plantation (A. van Harten, MZT 3066, PBI _ OON 9870 ), 1♀ (holotype) .

DISTRIBUTION: Yemen.

Khamisia atlit , new species

Figures 101–124 View FIGURES 101–115 View FIGURES 116–128. 116–124

TYPES: Male holotype and male paratype from Atlit, Haifa, Israel (Sept. 18–23, 2005; M. Vonshak), deposited in TAU ( PBI _OON 51500) .

ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.

DIAGNOSIS: Males resemble those of K. hayer but have a longer embolus with a smooth median portion (fig. 111); females differ from those of K. hayer and K. banisad in having narrow lateral genitalic processes without rounded tips (fig. 123).

MALE (PBI_OON 51500, figs. 101–116): Total length 0.95. ALE separated by less than their radius. Tip of endite rounded, with single, blunt-tipped seta. Embolus relatively long, straight.

FEMALE (PBI_OON 51501, figs. 117–124): Total length 1.07. Median genitalic process slightly expanded at tip, lateral processes narrow throughout their length.

OTHER MATERIAL EXAMINED: ISRAEL: Central: Gedera , Oct. 20–25, 2005 (M. Vonshak, TAU PBI _ OON 51501), 2♀ . Tel Aviv: Tel Aviv , Oct. 17–22, 2005 (M. Vonshak, TAU PBI _OON 51502), 2♂, 1♀ .

DISTRIBUTION: Israel.

Khamisia holmi , new species

Figures 125–143 View FIGURES 116–128. 116–124 View FIGURES 129–143

TYPE: Male holotype taken from litter in a ravine forest situated at an elevation of 600 m at a public campsite in Meru National Park , Isiolo, Kenya (Dec. 26, 1975; Å. Holm), deposited in UU (332, PBI _ OON 8532 ) .

ETYMOLOGY: The specific name is a patronym in honor of the collector of the holotype, the late Swedish arachnologist, Åke Holm.

DIAGNOSIS: Males can easily be recognized by their heavily sclerotized, distally excavated cheliceral fangs (figs. 138, 143).

MALE (PBI_OON 8532, figs. 125–143): Total length 1.09. ALE separated by less than their radius. Tip of endites with rounded protrusion bearing single, blunt-tipped seta. Embolus long, narrow, with ventral projection originating at about half its length.

FEMALE: Unknown.

OTHER MATERIAL EXAMINED: None.

DISTRIBUTION: Central Kenya.

Khamisina , new genus

TYPE SPECIES: Khamisina kivu , new species.

ETYMOLOGY: The generic name refers to the similarities to Khamisia , and is feminine in gender.

DIAGNOSIS: The bottlebrush-shaped setae on the cheliceral promargin (figs. 147, 204, 261) are diagnostic. These species differ from others with similarly enlarged sternal extensions in having a patterned abdomen (figs. 191, 251, 276) and a punctate sternum (figs. 149, 206, 236, 263, 280). The posterior median eyes resemble those of Longoonops in being relatively long and narrow (figs. 189, 249, 255, 276).

DESCRIPTION: Total length of males 1.1–1.3, of females 1.2–1.4. Cephalothorax and appendages yellow, without pattern, abdomen white except for pale yellow ventral scuta, with pattern (fig. 191). Cephalothorax: Carapace elongated hexagonal in dorsal view (figs. 144, 201), pars cephalica flat in lateral view, anteriorly narrowed to 0.49 times its maximum width or less, anterolateral corners with slightly sclerotized triangular projections, pars thoracica with rounded posterolateral corners, without depressions or radiating rows of pits, posterolateral edge without pits, posterior margin not bulging below posterior rim, posterolateral surface without spikes, surface of elevated portion of pars cephalica punctate, sides punctate (figs. 146, 203), fovea absent, lateral margin straight, rebordered, without denticles; plumose setae near posterior margin of pars thoracica absent; marginal, nonmarginal pars cephalica, pars thoracica setae dark, needlelike, scattered. Clypeus margin slightly rebordered (figs. 145, 202), straight in front view, sloping forward in lateral view, high, ALE separated from edge of carapace by more than their radius, slight median projection present; setae dark, needlelike. Chilum undivided. Eyes six, well developed, ALE largest, oval, PME relatively long, narrow (fig. 189), PLE oval; posterior eye row recurved from above, procurved from front; ALE separated by less than their radius, ALE-PLE separated by less than ALE radius, PME touching throughout most of their length, PLE-PME separated by less than PME radius. Sternum longer than wide, not fused to carapace, median concavity absent, with radial furrows between coxae I-II, II-III, III-IV, furrows smooth, radial furrow opposite coxae III absent, surface finely punctate (figs. 149, 206), without pits, microsculpture everywhere but middle, sickle-shaped structures absent, anterior margin with continuous transverse groove, posterior margin not extending posteriorly of coxae IV, anterior corner unmodified, lateral margin without infracoxal grooves, distance between coxae II and III greater than distance between coxae I and II or coxae III and IV (figs. 190, 196), extensions of precoxal triangles absent, lateral margins with rounded extensions between coxae, without posterior hump; setae sparse, dark, needlelike, evenly scattered, originating from surface; hair tufts absent. Chelicerae straight, anterior face unmodified; without teeth on promargin or retromargin; fangs without toothlike projections, directed medially, elongated, without prominent basal process, tip with elongated venom gland opening (figs. 148, 205, 262); setae dark, needlelike, evenly scattered; paturon inner margin with pairs of enlarged setae, distal region abruptly narrowed, posterior surface unmodified, promargin with row of bottlebrush-shaped setae (figs. 147, 204, 234), inner margin unmodified, laminate groove absent. Labium triangular, fused to sternum (figs. 150, 207), anterior margin indented at middle (figs. 154, 209), same as sternum in sclerotization; with six or more setae on anterior margin, subdistal portion with unmodified setae. Endites distally not excavated, same as sternum in sclerotization, serrula absent in males, present in females as single row of teeth (fig. 208), males with anteromedian tip elongated (figs. 195, 237, 264), posteromedian part of males sometimes with dorsally originating horn (figs. 150–153), flattened setae on tip serrated but without forklike tines. Labrum with flattened, triangular subdistal projection (figs. 155, 210). Female palp without claw or spines (figs. 211, 212) but with terminal nubbin presumably representing remnant of claw (figs. 285, 286); patella without prolateral row of ridges; tibia with three trichobothria, distalmost one situated near distal margin (figs. 213, 287); tarsus unmodified. Abdomen: Cylindrical, without long posterior extension, but females often with contents shrunken, leaving empty space under cuticle (fig. 255), rounded posteriorly, interscutal membrane rows of small sclerotized platelets absent posteriorly. Book lung covers large, ovoid, heavily sclerotized (fig. 198), without setae, anterolateral edge unmodified. Posterior spiracles connected by groove (figs. 165, 214). Pedicel tube short, unmodified, scutopedicel region unmodified, scutum not extending far dorsal of pedicel, plumose hairs, matted setae on anterior ventral abdomen in pedicel area, cuticular outgrowths near pedicel all absent. Dorsal scutum absent. Epigastric scutum weakly sclerotized, not surrounding pedicel, not protruding, small lateral sclerites absent, scuta of females without lateral joints. Postepigastric scutum weakly sclerotized, short, only around epigastric furrow (fig. 197), not fused to epigastric scutum, anterior margin unmodified, without posteriorly directed lateral apodemes. Spinneret scutum absent. Supraanal scutum absent. Abdominal setae dark, needlelike, epigastric area setae not basally thickened. Dense patch of setae anterior to spinnerets absent. Colulus present. Six spinnerets, anterior laterals bisegmented, basal segment with oblique membranous strip (figs. 166, 216), posterior medians unisegmented, posterior laterals bisegmented (figs. 167, 217), spigots scanned only in K. kivu , anterior laterals with one major ampullate gland spigot and two piriform gland spigots (figs. 168, 218), posterior medians with one spigot (figs. 169, 219), posterior laterals with two spigots (fig. 170, 220). Legs: Femora without subdistal constriction or subdistal transverse row of setae (figs. 171–174); femur IV not thickened, same size as femora I–III, patella plus tibia I shorter than carapace, tibia I unmodified, tibia IV specialized hairs on ventral apex, ventral scopula both absent, metatarsi I, II mesoapical comb absent, metatarsi III, IV weak ventral scopula absent. Leg spines absent. Tarsi I to IV without inferior claw. Superior claws of legs I–III with outer row of three or four large, blunt-tipped teeth (figs. 180–182, 225–227), inner row with distal series of closely spaced teeth (figs. 176–178, 221–223), those of leg IV apparently without inner tooth row, with only median row of short, proximally situated teeth (figs. 179, 183, 224, 228). Tarsal organs with two long, raised receptors, distal receptor bifid, pore receptor possibly present on anterior legs (figs. 184–188, 229–233). Trichobothrial base with distinct distal portion marked by ridges running perpendicular to other cuticular ridges (fig. 175). Genitalia: Male epigastric region with sperm pore not visible; furrow without Ω-shaped insertions, without setae. Male palp normal size, not strongly sclerotized, right and left palps mirror images; trochanter normal size, unmodified; femur normal size, two or more times as long as trochanter, without posteriorly rounded lateral dilation, attaching to patella basally; patella shorter than femur, not enlarged, without prolateral row of ridges, setae unmodified; tibia with three trichobothria, distalmost one situated near distal margin (figs. 156, 241, 268); cymbium ovoid in dorsal view, completely fused with bulb, no seam visible (figs. 157–160), not extending beyond distal tip of bulb, plumose setae, stout setae, distal patch of setae all absent; bulb 1–1.5 times as long as cymbium, stout, spherical; embolus light, without prolateral excavation, wide, tubelike (figs. 192–194), with subdistal prong (figs. 161–164). Females with two internesting V-shaped sclerites (figs. 199, 200), anterior sclerite with posteriorly directed glands (fig. 215).

DISTRIBUTION: Tropical Africa ( Nigeria to Kenya).

TAU

Tel-Aviv University

UU

University of Uppsala

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Oonopidae

Genus

Khamisia

Loc

Khamisia banisad Saaristo and van Harten

Platnick, Norman I. & Berniker, Lily 2015
2015
Loc

Khamisia banisad

Saaristo and van Harten 2006: 136
2006
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