Eupoecilia

Zhang, Xu & Li, Houhun, 2008, Review of the Chinese species of Eupoecilia Stephens (Lepidoptera: Tortricidae: Cochylini), Zootaxa 1692, pp. 55-68 : 64-67

Key to Eupoecilia species of the Oriental and Palaearctic regions

1 Male ............................................................................................................................................................ 2

– Female ....................................................................................................................................................... 28

2 Median process of transtilla broader than width of socius; socius well sclerotized ................................... 3

– Median process of transtilla narrower than or equal to width of socius; socius weakly sclerotized ........ 21

3 Aedeagus with comblike distal lobes projecting dorsally .......................................................................... 4

– Aedeagus without such comblike lobes ...................................................................................................... 6

4 Socius distinctly dentate at base ( Razowski, 1968: fig. 22) ...................................... E. dentana Razowski

– Socius not dentate at base ........................................................................................................................... 5

5 Vesica with one large cornutus almost equal in length to aedeagus ( Razowski, 1968: fig. 25) .................... ............................................................................................................................ E. amphimnesta (Meyrick)

– Vesica with one large cornutus about half the length of aedeagus ( Razowski, 1968: fig. 21) ...................... ................................................................................................................................... E. armifera Razowski

6 Sacculus with spines ................................................................................................................................... 7

– Sacculus without spines ............................................................................................................................ 12

7 Spines on sacculus thick, about 1/2 width of socius ................................................................................... 8

– Spines on sacculus slender, about 1/5 width of socius ............................................................................... 9

8 Vesica with a large cornutus at the base and one at the middle; costa of valva gently concave ( Diakonoff, 1984: figs. 13, 14) ...................................................................................................... E. coniopa Diakonoff

– Vesica without large cornutus at base and middle; costa of valva nearly straight ( Diakonoff, 1983: figs. 5, 6) ............................................................................................................................... E. anebrica Diakonoff

9 Aedeagus with distal cornuti having flat base .......................................................................................... 10

– Aedeagus with distal cornuti lacking flat base (fig. 2b) ........................................ E. ambiguella (Hübner)

10 Vesica with three strong cornuti near middle; central part of transtilla narrow, 2.0-2.5 times width of socius ........................................................................................................................................................ 11

– Vesica with group of more than ten slender cornuti near middle; central part of transtilla broad, 3.5 times

width of socius ( Razowski, 1968: figs. 4, 5) ................................................... E. tenggerensis (Diakonoff) 11 Cornuti having flat base; central part of transtilla twice width of socius ( Razowski, 1968: figs. 7, 8) ......... ................................................................................................................................ E. sumbana (Diakonoff)

– Cornuti lacking flat base; central part of transtilla 2.5 times width of socius ( Razowski, 1968: figs. 10, 11) .......................................................................................................................... “ E. reliquatrix (Meyrick) ”.

12 Valva broadly subquadrate ........................................................................................................................ 13

– Valva subtriangular or oblong ................................................................................................................... 14

13 Vesica with the largest cornutus about 4/5 of length of aedeagus ( Diakonoff, 1983: fig. 4) ......................... ................................................................................................................................ E. sumatrana Diakonoff

– Vesica with the largest cornutus about 1/6 of length of aedeagus (fig. 3b) ..................... E. inouei Kawabe

14 Valva subtriangular ................................................................................................................................... 15

– Valva oblong ............................................................................................................................................. 20

15 Vesica with distal cornuti .......................................................................................................................... 16

– Vesica without distal cornuti ..................................................................................................................... 19

16 Vesica with groups of small cornuti at base and near middle ( Diakonoff, 1984: fig. 10) ............................. ............................................................................................................................... E. engelinae (Diakonoff)

– Vesica with one large cornutus at base or middle ..................................................................................... 17

17 Vesica with cornutus at base ..................................................................................................................... 18

– Vesica with cornutus at middle ( Razowski, 2005: fig. 4) ....................................... E. yubariana Razowski

18 Aedeagus twice length of valva, with an apical projection (fig. 3d). E. sanguisorbana (Herrich-Schäffer)

– Aedeagus 1.25 times length of valva, without apical projection (fig. 2c) ............... E. angustana (Hübner)

19 Vesica with three strong cornuti; central part of transtilla narrow, twice width of socius ( Razowski, 2005: figs. 5, 6) ........................................................................................................................... E. lata Razowski

– Vesica with one strong cornutus; central part of transtilla broad, 3.5 times width of socius ( Razowski, 1968: figs. 16, 17) ....................................................................................................... E. crocina Razowski

20 Vesica with one straight cornutus extending from base to near middle, distal cornuti capitately arranged (fig. 3a) ..................................................................................................................... E. citrinana Razowski

– Vesica with one bent cornutus extending from base to apex, distal cornuti absent ( Razowski, 1970a: Tafel 76: fig. 171) ................................................................................................................ E. cebrana (Hübner)

21 Central part of transtilla narrower than width of socius ........................................................................... 22

– Central part of transtilla about equal to width of socius ........................................................................... 24

22 Cucullus as wide as base of valva ( Razowski, 1968: fig. 41) .............................. E. charixantha (Meyrick)

– Cucullus narrower than base of valva ....................................................................................................... 23

23 Vesica with one large cornutus (fig. 3c) .................................................................... E. kobeana Razowski

– Vesica lacking large cornutus ( Razowski, 1968: fig. 45) ....................................... E. eucalypta (Meyrick)

24 Sacculus bearing prominence with denticulate ventral edge .................................................................... 25

– Sacculus lacking such prominence ........................................................................................................... 26

25 Vesica with four strong distal cornuti, with a subapical and a second spiny projection; costa of valva nearly straight ( Diakonoff, 1984: figs. 11, 12) ....................................................... E. wegneri (Diakonoff)

– Vesica with five strong distal cornuti, with one subapical projection only; costa of valva gently concave (fig. 2a) ..................................................................................................................... E. quinaspinalis sp. n.

26 Sacculus with small spines posteriorly; aedeagus broadly parallel-sided, vesica with a cornutus about 2/3 length of aedeagus ( Razowski, 1968: figs. 35, 36) ................................................. E. ochrotona Razowski

– Sacculus without spines; aedeagus elongate pear-shaped, vesica with a cornutus about half length of aede- agus ............................................................................................................................................................ 27

27 Cucullus oblong ( Diakonoff, 1982: fig. 1) ........................................................... E. anisoneura Diakonoff

– Cucullus subtriangular ( Diakonoff, 1982: pl. I, fig. 1) .............................................. E. cracens Diakonoff

28 Ductus bursae broadened posteriorly ........................................................................................................ 29

– Ductus bursae not broadened posteriorly .................................................................................................. 32

29 Corpus bursae entirely spined ( Diakonoff, 1984: fig. 15) ......................................... E. coniopa Diakonoff

– Corpus bursae partially spined .................................................................................................................. 30

30 Corpus bursae with long spines forming a “star” at middle ..................................................................... 31

– Corpus bursae without such spines at middle (fig. 4b) .......................................... E. ambiguella (Hübner)

31 Ductus bursae broad, about 2/3 width of corpus bursae; corpus bursae with row of long spines laterally ( Razowski, 1968: fig. 6) .................................................................................. E. tenggerensis (Diakonoff)

– Ductus bursae narrow, about 1/3 width of corpus bursae; corpus bursae with spines irregularly scattered ( Razowski, 1968: fig. 12) ...................................................................................... E. reliquatrix (Meyrick)

32 Ductus bursae length less than twice its greatest width ............................................................................ 33

– Ductus bursae length greater than twice its greatest width ....................................................................... 40

33 Sterigma with denticulate edge posteriorly .............................................................................................. 34

– Sterigma without denticulate edge ............................................................................................................ 36

34 Ductus bursae narrow, about 1/2 width of corpus bursae; corpus bursae with spines forming radiating arc at the anterior edge .................................................................................................................................... 35

– Ductus bursae broad, about 3/4 width of corpus bursae; corpus bursae with spines not forming radiating arc at the anterior edge ( Razowski, 1968: fig. 27) .................................................. E. wegneri (Diakonoff)

35 Ductus seminalis originating from anterior 1/3 of ductus bursae (fig. 4a) ............... E. quinaspinalis sp. n.

– Ductus seminalis originating from proximal anterior portion of ductus bursae ( Diakonoff, 1984: fig. 16). ....................................................................................................................................... E. thalia Diakonoff

36 Ductus bursae entirely membranous ......................................................................................................... 37

– Ductus bursae partially sclerotized ........................................................................................................... 38

37 Sterigma protruding at middle on anterior margin; ductus bursae 1/2 length of corpus bursae ( Razowski, 1968: fig. 26) ..................................................................................................... E. amphimnesta (Meyrick)

– Sterigma concave at middle on posterior margin; ductus bursae 1/3 length of corpus bursae ( Diakonoff, 1971: fig. 15) ..................................................................................................... E. dynodesma (Diakonoff)

38 Apophyses about 1/3 width of papilla analis ( Razowski, 1970a: Tafel 141: fig. 171) E. cebrana (Hübner)

– Apophyses about 1/6 width of papilla analis ............................................................................................ 39

39 Ductus bursae about 2/3 sclerotized laterally and posteriorly ( Razowski, 1970a: Tafel 141: fig. 172) ........ .................................................................................................................................. E. angustana (Hübner)

– Ductus bursae about 1/2 sclerotized anteriorly ( Razowski, 1968: fig. 9) .............. E. sumbana (Diakonoff)

40 Corpus bursae with spines forming radiating arc at the anterior edge (fig. 4d) .............. E. inouei Kawabe

– Corpus bursae with spines not forming radiating arc ............................................................................... 41

41 Antrum with minute spines posteriorly .................................................................................................... 42

– Antrum without spines .............................................................................................................................. 43

42 Ductus bursae as long as corpus bursae; corpus bursae entirely spined ( Razowski, 1968: fig. 43) .............. .............................................................................................................................. E. charixantha (Meyrick)

– Ductus bursae twice length of corpus bursae; corpus bursae lacking spines medially ( Razowski, 1968: fig. 46) ........................................................................................................................... E. eucalypta (Meyrick)

43 Corpus bursae entirely spined ................................................................................................................... 44

– Corpus bursae partially spined................................................................................................................... 45

44 Ductus bursae narrow, 1/4 width of corpus bursae; corpus bursae oblong (fig. 4e) .. E. kobeana Razowski

– Ductus bursae broad, 1/3 width of corpus bursae; corpus bursae round ( Razowski, 2005: fig. 7) ............... ................................................................................................................................. E. yubariana Razowski

45 Corpus bursae with three spines ( Razowski, 1968: fig. 18) ....................................... E. crocina Razowski

– Corpus bursae with more than ten spines ................................................................................................. 46

46 Ductus bursae with anterior portion as broad as posterior portion ........................................................... 47

– Ductus bursae with anterior portion broader than posterior portion ( Razowski, 1968: fig. 14) .................... ........................................................................................................................................... E. lata Razowski

47 Ductus bursae nearly as long as corpus bursae (fig. 4f) ................................................................................ ............................................................................................................ E. sanguisorbana (Herrich-Schäffer)

– Ductus bursae longer than corpus bursae (fig. 4c) .................................................... E. citrinana Razowski