Khamul tolkeini Gates

Gates, M. W., 2008, Description of Khamul, gen. n. (Hymenoptera: Chalcidoidea: Eurytomidae), with a hypothesis of its phylogenetic placement, Zootaxa 1898 (1), pp. 1-33: 31-32

publication ID 10.11646/zootaxa.1898.1.1

persistent identifier

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scientific name

Khamul tolkeini Gates

n. sp.

Khamul tolkeini Gates   , n. sp.

( Figs. 54–57 View FIGURES 52–59 )

Etymology. tolkeini   (Latinized, noun) = genitive, singular, masculine; named in honor of J. R. R. Tolkein for his profound impact on the fantasy literature genre.

Diagnosis and identification. Tegula brown, hind tibia black. Preorbital carinae robustly protruding ventrally, delimiting a reniform subocular fovea ( Fig. 54 View FIGURES 52–59 ), furthermore, carinae expanded dorsally as triangular processes between lateral ocellus and eye ( Fig. 54 View FIGURES 52–59 ). Propodeal median channel distinctly delimited, composed of transverse reniform punctae, bordered laterally by smaller punctae and irregular rugosity (Fig. 64).

Description. Female Holotype. Length 3.3 mm. Head, body, and coxae black. Scape black; pedicel black basally, brown apically; flagellum dark brown. Femora, metatibia, metatarsomeres 1+2 black; pro- and mesotibiae and tarsomeres, extreme apex of metafemur, and metatarsomeres 3–5 brown ( Fig. 55 View FIGURES 52–59 ).

Head. 0.78X as broad as high; 1.1X wider than pronotum; HTE:msp 1.81. POL 2.5X as long as OOL. Width of oral fossa 0.42X width of head. Preorbital carina of uniform width, widening beneath eye, reniform subocular fovea present, carina produced as triangular tooth at vertex ( Fig. 54 View FIGURES 52–59 ). Scape 3.0X as long as broad. Antennal segment relative measurements 39:7:1:40:31:30:20:35:21:35. Clava 2.33X as long as broad.

Mesosoma. Dorsal pronotum 1.96X as broad as long. Mesoscutal midlobe 1.1X as broad as long. Scutellum 0.71X as broad as long; broadly convex dorsally, scutellar boss present, ~2.0X larger than any adjacent umbilicate puncta and its surrounding border combined. Dorsellum disc composed of a central -shaped structure connected laterally to submedian carinae ( Fig. 56 View FIGURES 52–59 ). Propodeal median channel composed of transverse reniform punctae, channel bordered laterally by smaller punctae and irregular rugosity ( Fig. 56 View FIGURES 52–59 ). Relative measurements marginal:postmarginal:stigmal veins as 13:35:32; stigmal vein evenly arcuate ( Fig. 57 View FIGURES 52–59 ); infuscation darkest along venation, fading toward disc margins.

Metasoma. Petiole 3.0X as broad as long in dorsal view; fine transverse carina anteriorly, protruberant lat- erally; bicarinate ventrally, defining triangular fovea, produced in lateral view. Gastral terga measurements in dorsal view on median line as 15:25:45:30:10:2, syntergum not visible in dorsal view.

Male. Unknown.

Variation. The type series exhibits very little variation.

Type material (1♀). Holotype: ♀ (held in trust at USNM); ECUADOR: Orellana: Tiputini Biodiversity Station , 216m, 0037’55” S 76°08’39” W, 9.ii.1999, Lot 2016, Trans. 2, T. Erwin et al. Canopy fogging bare leaves, some with bryophytic/lichenous coat; Restrictions Apply NMNH-DCB / EPN, Agreement 39   . Paratype ♀: PERU: Loreto, Explomapo Camp, Riio Napo, Rio Suscari ,, 03° 15’S 072° 55’W, Lot #382, T. L. Erwin, hand. coll. (♀ USNM) GoogleMaps   .


Smithsonian Institution, National Museum of Natural History


Tavera, Department of Geology and Geophysics


Escuela Polytecnica Nacional