Khamul gothmogi Gates

Gates, M. W., 2008, Description of Khamul, gen. n. (Hymenoptera: Chalcidoidea: Eurytomidae), with a hypothesis of its phylogenetic placement, Zootaxa 1898 (1), pp. 1-33: 29-30

publication ID 10.11646/zootaxa.1898.1.1

persistent identifier

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scientific name

Khamul gothmogi Gates

n. sp.

Khamul gothmogi Gates   , n. sp.

( Figs. 31 View FIGURES 26–33 , 65–67)

Etymology: gothmogi   (Latinized, noun) = genitive singular, masculine, named in honor of Gothmog, Lord of the Balrogs, High Captain of Angband ( Tolkein 1977).

Diagnosis and identification: Tegula black, apical mandibular tooth as an equilateral triangular and curved. Preorbital carina is finely produced ventrally but does not delimiti a reniform subocular fovea; carinae expanded into a rounded process between lateral ocellus and eye (Fig. 65). Scutellum convex in lateral view (Fig. 66) as opposed flat in K. erwini   ( Fig. 37 View FIGURES 36–43 ). Propodeal median channel is distinctly delimited, particularly anteriorly (Fig. 67).

Description: Female holotype. Length 3.3 mm. Head, scape, tegula, body, coxae, and basal 7/8 metafemur black. Basal third pedicel, flagellum, and basal 2/3 pro- and mesofemur brown. Apices femora, tibiae, tarsi pale golden. Pretarsus brown. Fore wing infuscate, venation brown.

Head. 1.41X as broad as high; 1.1X wider than pronotum; HTE:msp 2.0. POL 3.0X as long as OOL. Width of oral fossa 0.40X width of head. Clypeus not separated from supraclypeal area by carina. Preorbital carina of uniform width beneath eye, reniform subocular fovea absent, carina produced as blunt lobe at vertex. Scape 3.33X as long as broad. Antennal segment relative measurements 34:6:1:35:25:26:23:22:20:35. Clava 2.33X as long as broad.

Mesosoma. Dorsal pronotum 2.0X as broad as long. Mesoscutal midlobe approximately as broad as long. Scutellum 0.67X as broad as long; broadly convex dorsally, scutellar boss present, subequal in dimensions to three adjacent umbilicate punctae and surrounding border combined, distinctly convex in lateral view (Fig. 66). Dorsellum disc composed of a central -shaped structure, depressed dorsally, connected laterally to submedian carinae (cf. Fig. 64); posterior invagination entire. Propodeal median channel distinct, delimited laterally by carinae, composed of rectangular foveae (Fig. 67), channel bordered laterally by numerous smaller punctae. Relative measurements marginal:postmarginal:stigmal veins as 21:37:33; stigmal vein arcuate.

Metasoma. Petiole 3.0X as broad as long in dorsal view; fine transverse carina anteriorly, protruberant laterally; bicarinate ventrally, defining triangular fovea, produced in lateral view. Gastral terga measurements in dorsal view on median line as 20:30:33:23:15:6, syntergum not visible in dorsal view.

Male. Unknown.

Variation. The paratype from Waorani has a stigmal vein that is slightly bent (angulate) rather than smoothly arcuate. Also, the preorbital protuberance of the specimen is slightly smaller and narrower than that of the holotype. The area occupied by the scutellar boss also is slightly variable among members of the type series, ranging from 1–3 foveal equivalents.

Type material ( 3♀, held in trust at USNM). Holotype ♀; ECUADOR: Orellana: Tiputini Biodiversity Station , 216m, 0037’55” S 76°08’39” W, 9.ii.1999, Lot 2017, Trans 2, T. Erwin et al. Canopy fogging bare leaves, some with bryophytic/lichenous coat; Restrictions Apply NMNH-DCB / EPN, Agreement 39. Paratypes: 3♀ (2♀ held in trust at USNM); same as holotype, but 9.ii.99, Lot 2000, Trans 1; ECUADOR: Napo: Reserva Etnica Waorani, Transect Ent., 1km S. Okone Gare Camp, 0039’10” S 76°26’00” W, 220m,  

5.ii.1996, T   . Erwin et al. Canopy fogging, t3..3, #1423; Restrictions Apply NMNH-DCB / EPN, Agreement 39; PERU: Loreto, Explomapo Camp, Riio Napo, Rio Suscari   ,, 03 15’S 072° 55’W, Lot #382, T GoogleMaps   . L. Erwin, hand. coll. (♀ USNM)   .


Smithsonian Institution, National Museum of Natural History


Tavera, Department of Geology and Geophysics


Escuela Polytecnica Nacional