Manota panda, Hippa, Heikki & Kurina, Olavi, 2013

Hippa, Heikki & Kurina, Olavi, 2013, New species and new records of Manota Williston (Diptera: Mycetophilidae) in the Neotropical region, Zootaxa 3686 (2), pp. 101-139 : 120-122

Manota panda sp. n.

Figs. 11 A–D

Male. Colour. Head brown, face paler brown. Antenna pale brown. Clypeus and mouthparts pale yellowish. Thorax brown, scutum and scutellum seem darker medially. Legs yellow, femur 2 with large infuscated patch basoventrally, femur 3 with more intensive infuscated patch on basoventral third. Wing pale greyish; haltere pale brown with darker brown knob. Abdomen pale brown, sternites paler than tergites. All setosity pale, yellowish or brownish, the thicker setae seeming darker than the finer ones. Head. Antennal flagellomere 4 seen in oblique ventral view in the single specimen, apparently similar to Fig. 4 A ( M. appendiculata ). Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 3 apically curved sensilla; palpomere 4 with parasegment; palpomere 5 ca. 1.3 times longer than palpomere 4. Number of strong postocular setae 9–11. Thorax. Anepisternum setose, with 73 setae; anterior basalare non-setose; preepisternum 2 non-setose; laterotergite nonsetose; episternum 3 setose, with 15 setae. Legs. Mid and hind tibial organs absent. Wing. R1 meeting C well on the basal half of the costal margin; the sclerotized part of M2 not extending to the level of the tip of R1; wing length 1.8 mm. Hypopygium. Figs. 11 A–D: Sternite 9 laterally fused with gonocoxa, the posterior margin short, transverse, extending to the middle of gonocoxa, anterior margin deeply incised, the setae similar to the adjacent ventral setae of gonocoxa. Ventral mesial margin of gonocoxa wavy, the ventral setae unmodified; posterolateral part is drawn into a long posteriorly widening lobe, the setae of which are arising from small ridges. Parastylar lobe fused with gonocoxa, apparently the seta drawn in Fig. 11 B on the mesial margin of gonocoxa belongs to parastylar lobe. No paraapodemal lobe identifiable. Dorsal mesial margin of gonocoxa simple, poorly visible in the specimen studied; on posterior part, at the juxtagonostylar megasetae, with a low ridge bearing an aggregation of setae. Number of juxtagonostylar setae two, the ventral one flattened and slightly expanded megaseta, the dorsal one complicated with expanded medial part, both of the megasetae arising from a common basal body which is shorter than the megasetae. Anteriorly from the juxtagonostylar megasetae there is a plate-like lobe bearing a strong megaseta. Between this seta and the juxtagonostylar setae exist on more ventral level as a subtriangular membranous lobe with two blunted setae. Posteriorly from the juxtagonostylar megasetae there is a group of ca. 4 strong setae deviating from the other setosity. Otherwise the dorsal setae of gonocoxa similar to those on the ventral side. Gonostylus flat, bilobed, the apicomesial lobe with two long setae, the apicolateral lobe with a stronger curved short seta. In the mount of the holotype there is a lobe on the dorsal side of gonostylus bearing a group of long setae at apex and a few short setae at the base. It seems that this lobe is a part of the gonostylus, but we are not sure about it. Aedeagus subtriangular, the lateral sides straight except for a small subtriangular lobe, the apex curved ventrad. Hypoproct extending posteriorly to middle of gonostyli, the number of ventral setae (sternite 10) ca. 30 on each half. Cerci mesially separate.

Female. Unknown.

Discussion. The non-setose anterior basalare, the non-setose laterotergite, sternite 9 fused laterally with the gonocoxa, the lack of distinct sclerotized parastylar lobe, the well-developed posterolateral lobe on the gonocoxa, the strong oblique megaseta anteriorly from the juxtagonostylar megasetae, and the group of strong setae on a lobe posteriorly from the juxtagonostylar megasetae make Manota panda reminiscent of M. costaricensis Jaschhof & Hippa , M. fraterna Jaschhof & Hippa and M. parva Jaschhof & Hippa. Also , the gonostylus is similarly subtriangular with a concave apex (if the dorsal lobe mentioned above in the description does not belong to gonostylus). This lobe is similar to a lobe just anterior from the juxtagonostylar megasetae in these three species but cannot be localized in the mount of the holotype of M. panda . Manota panda is distinguished from the other similar species by having a long setose crest at the dorsal mesial margin of the gonocoxa, at the level of the juxtagonostylar megasetae and by having the posterior margin of sternite 9 very narrow, about half of the length of its setae instead of being ca. twice the length of the setae and by having slightly shorter and apically broadening posterolateral lobe on the gonocoxa. Like M. parva , M. panda has a non-setose ventral surface on the gonostylus.

Etymology. The name is Latin, panda , crooked, bent, referring to the strongly bent apicolateral lobe on the gonostylus.

Types. Holotype. Male, ARGENTINA, Balneario San Juan, 10 km ENE of Oberá Missiones, 13.xi.2004, T. Saigusa col. (on slide, in KUM).

Figures

FIGURE 4. Manota appendiculata sp. n. (A paratype from French Guyana, B, C, D holotype). A. Antennal flagellomere 4, lateral view. B. Hypopygium, dorsal view. C. Hypopygium, ventral view. D. Aedeagus and hypoproct, ventral view. E. Gonostylus, dorsal view. F. Juxtagonostylar megasetae, dorsal view. Scale 0.1 mm.

FIGURE 11. Manota panda sp. n. (holotype). A. Hypopygium, dorsal view. B. Hypopygium, ventral view. C. Gonostylus, ventral view. D. Aedeagus and hypoproct, ventral view. Scale 0.1 mm.