Dipogon (Stigmatodipogon) budrisi ( Loktionov & Lelej, 2014 ), 2018

Shimizu, Akira, Lelej, Arkady S., Loktionov, Valery M., Nishimoto, Yutaka & Endo, Tomoji, 2018, Revision of the subgenus Stigmatodipogon Ishikawa of the genus Dipogon Fox (Hymenoptera: Pompilidae: Pepsinae), Zootaxa 4514 (1), pp. 1-22 : 3-7

1. Dipogon (Stigmatodipogon) budrisi ( Loktionov & Lelej, 2014) , comb. nov.

( Figs 1 , 2 )

Dipogon (Stigmatodipogon) petiolatus: Lelej 2005: 126 , ♀ ♂, misidentification.

Stigmatodipogon petiolatus: Loktionov & Lelej 2012b: 317 , ♂, misidentification.

Stigmatodipogon budrisi Loktionov & Lelej, 2014: 158 , 160, ♀ ♂ (holotype ♂, Russia, Kuril Islands, Kunashir Island, 2 km NE Tretyakovo, 10.viii.1986 (E. Budris) [IBSS], examined).

Diagnosis. FEMALE. This species differs from other species by having the gena in dorsal view rather developed, gently and roundly receding posteriorly in both sexes ( Figs 1A , 2A ); the clypeus in large part, the mandible, scape, pedicel, and legs yellowish-brown ( Figs 1B, 1C, 1D ); and the frons impunctate with minute setiferous pores, interspaces almost smooth and strongly polished in the female ( Fig. 1B ); and having metasomal tergum I barely petiolate ( Fig. 2B ) and the transverse groove on metasomal sternum II almost straight in the male.

Redescription. FEMALE (holotype). Length: body 4.2 mm; fore wing 3.9 mm. Body black and polished ( Fig. 1C ). Following yellowish-brown: clypeus (dark-brown basilaterally) ( Fig. 1B ), mandible (reddish-brown apically), palpi, scape ( Figs 1C, 1D ), pedicel and all legs (coxae laterally and dorsally and tarsomeres V distally castaneous) ( Fig. 1C ). Following bright-brown: flagellum (darker above than below) ( Fig. 1C ), lateral margin of pronotum, tegula ( Figs 1A, 1E ), posteromedian spots of metasomal terga II, III and of sterna I–V, and terga IV posteriorly. Wings hyaline ( Figs 1F, 1G ); fore wing fasciae indistinctly defined.

Body with short, greyish- to yellowish-white pubescence. Not setose; vertex and dorsa of pro- and mesonota with a few pale setae; clypeus preapically and mandible with long yellowish-brown bristles arising from somewhat large pores; metasomal sterna with sparse brown bristles (terga V, VI and sternum VI missing).

Head weakly alutaceous and strongly polished, with minute setiferous pores ( Fig. 1B ). Dorsum of pronotum impunctate and alutaceous. Dorsum of mesoscutum densely punctate and strongly alutaceous. Scutellum and metanotum weakly punctate, interspaces smooth. Mesepisternum and lower metapleuron strongly alutaceous. Side of metanotum rather regularly and strongly striate. Propodeum irregularly punctate and strongly alutaceous.

Head width 1.1 × length, 1.4 × pronotal dorsum width. Vertex in frontal view moderately convex ( Fig. 1B ). Inner orbits convergent distinctly above, barely below. UID: MID: LID = 7.3: 10: 9.0. MID 0. 63 × head width. Frons broad, in profile evenly convex, protruding anteriorly, far exceeding level of clypeus ( Fig. 1D ); frontal line sharply impressed only below. Gena width in lateral view 0.7 × eye width. Ocellar triangle acute-angled, scarcely raised. POL: OOL = 1: 1. Clypeus slightly narrower than LID ( Fig. 1B ), its width 2.4 × length, convex medially; apicolateral corner rounded; apical rim slightly depressed, set off from main surface; apical margin feebly convex. Malar space short. Flagellomere I length 1.3 × flagellomere II length, 5.2 × its width, 0.76 × UID.

Pronotum with anterior declivity vertical ( Figs 1C, 1H ), feebly differentiated from dorsum; shoulder slightly bulging ( Figs 1A, 1E ); posterior margin with obtuse angulation medially. Discs of scutellum and metanotum distinctly convex above level of mesoscutum ( Fig. 1H ). Metapostnotum length 0.1 × metanotum length at midline. Propodeum length 1.0 × width; dorsum in profile feebly convex, merging into declivity, with median groove.

Mid and hind tibiae with a few minute spines laterally. Longer spur of hind tibia 0.5 × hind tarsomere I length. Hind tarsomeres I–IV thin; tarsomere V slightly enlarged, its length 1.3 × tarsomere IV length, its width 1.2 × tarsomere IV width. Tarsal claws with small acute tooth preapically.

Fore wing shown in Fig. 1F . Pterostigma length 1.9 × SMC2 length, its width 0.78 × SMC2 width. SMC2 length 0.71 × width, narrowed on vein Rs by 0.72 × its own length on vein M, receiving crossvein 1m-cu at basal 0.44. SMC3 length 1.3 × width, 1.1 × SMC2 length on vein M, 1.0 × SMC2 length on vein Rs, narrowed on vein Rs by 0.65 × its own length on vein M, receiving crossvein 2m-cu at basal 0.43. Crossvein 2rs-m barely curved. Crossvein 3rs-m arcuate. Hind wing crossvein cu-a forming obtuse angle with vein A, short and slightly sinuate ( Fig. 1G ).

Metasomal tergum I distinctly petiolate ( Fig. 1I ). Tergum II width 0.86 × head width.

Redescription. MALE (paratype). Length: body 3.9 mm; fore wing 3.8 mm. Body black ( Fig. 2C ). Mandible dark brown on basal half, yellowish-brown on apical half, reddish-brown apically ( Fig. 2D ). Palpi and tegula light brown ( Figs 2A, 2E ). Following ferruginous: clypeus apically, antenna below, lateral margin of pronotum, posterior margins of metasomal terga, laterotergum, sternum I mostly and sternum II basally. Legs mostly dark brown; tarsi darker distally ( Fig. 2C ); fore and mid tibiae yellowish-brown below; fore femur and tarsus, mid basitarsus, and hind tibia below bright-brown.

Setae sparse; a pair of setae on vertex close to inner orbits and setae on clypeus, mandible, gena, postgena, propleuron, mesosternum and fore coxa comparatively long.

Head width in frontal view 1.2 × length. Inner orbits convergent both above and below ( Fig. 2D ). UID: MID: LID = 8.3: 10: 8.8. MID 0.61 × head width. Lower frons in profile strongly protruding anteriorly as in female ( Fig. 2E ). Ocellar triangle right-angled ( Fig. 2A ). POL: OOL = 1: 0.78. Clypeus barely narrower than LID ( Fig. 2D ), its width 2.3 × length, spherically convex medially; apical rim not depressed, very narrow, smooth and polished; apical margin almost straight. Gena in dorsal view well developed as in female ( Fig. 2A ), its width in profile 0.7 × eye width ( Fig. 2E ). Flagellomere I length 1.0 × flagellomere II length, 2.7 × its width, and 0.61 × UID.

Pronotum with shoulder roundly receding anteriorly ( Fig. 2A ). Metapostnotum length 0.2 × metanotum length at midline, triangularly emarginate posteromedially. Propodeum as long as broad, in profile more gently convex than in female ( Fig. 2C ).

Wings shown in Figs 2F and 2G . Pterostigma length 2.4 × SMC2 length, its width 0.85 × SMC2 width. SMC2 length 1.3 × width, narrowed on vein Rs by 0.82 × its own length on vein M, receiving crossvein 1m-cu at basal 0.56; SMC3 length 1.2 × width, 1.1 × SMC2 length on vein M, 0.64 × SMC2 length on vein Rs, narrowed on vein Rs by 0.49 × its own length on vein M, receiving crossvein 2m-cu at basal 0.32. Crossvein 3rs-m strongly sinuate.

Longer spur of hind tibia 0.60 × hind tarsomere I length. Tarsal claws with minute tooth preapically.

A transverse groove on sternum II close to posterior margin of tergum I. A pair of lateral hooks on sternum VI tapering and directed outward; posterior margin of sternum VI between hooks triangularly emarginate. Exposed portion of SGP ( Fig. 2H ) triangularly produced on sides with dense setae subapically and apically; apical margin roundly convex. Genitalia ( Fig. 2I ) with paramere barely extending beyond apex of parapenial lobe.

Material examined. Paratype. RUSSIA. Kuril Islands , Kunashir Island: 1♂, 2 km NE Tretyakovo, 10.viii.1986 (E. Budris) [ IBSS] .

Distribution. Russia (Far East: Kunashir Island).

Biology. Unknown.

Figures

FIGURE 1. Dipogon (Stigmatodipogon) budrisi, holotype, ♀. A. Head and pronotum, dorsal view. B. Head, frontal view. C. Whole body, lateral view. D. Head and pronotum, lateral view. E. Head, pro-, meso- and metanota, dorsal view. F. Fore wing. G. Hind wing. H. Mesosoma anteriorly, lateral view. I. Metasomal tergum I basally, dorsal view. Scale bars: 1 mm for C, 0.1 mm for I, 0.5 mm for others.

FIGURE 2. Dipogon (Stigmatodipogon) budrisi, paratype, ♂. A. Head, pronotum and scutum, dorsal view. B. Metasomal tergum I basally, dorsal view. C. Whole body, lateral view. D. Head, frontal view. E. Head and pronotum, lateral view. F. Fore wing. G. Hind wing. H. Metasomal sternum VII and SGP, ventral view. I. Genitalia, ventral view. Scale bars: 1 mm for C, 0.1 mm for B, 0.2 mm for H, I, 0.5 mm for others (H, I from Loktionov & Lelej 2014).