Paramblynotus costaricanus, Liu, Ronquist & Nordlander, 2007

Liu, Zhiwei, Ronquist, Fredrik & Nordlander, Göran, 2007, The Cynipoid Genus Paramblynotus: Revision, Phylogeny, And Historical Biogeography (Hymenoptera: Liopteridae), Bulletin of the American Museum of Natural History 2007 (304), pp. 1-151 : 83-84

Paramblynotus costaricanus , new species

FEMALE: Length 4.5 mm. Head and pronotum yellow brown; mesonotum black; rest of the body, antenna, and legs brown to dark brown. Forewing with a large macula covering marginal cell, distal third of first submarginal cell and basal third of second submarginal cell, a narrow band along interior side of basal vein; both wings without a wide band along outer margin.

Antenna 13-segmented; flagellum filiform. Vertex glabrate except a posterior row of foveae along border between vertex and occiput. Eye prominent, distinctly extended laterally beyond outer margin of gena. Ocellar plate distinctly raised, glabrate anterior to anterior ocellus and foveate-reticulate posteriorly, and defined laterally by distinct carina. Median frontal carina not extended to lower face and distinctly raised into a prominent, laminate, and apically flattened process. Upper face coriarious laterally; antennal scrobe glabrous with setigerous punctures anteriorly; antennal scrobe deeply depressed and defined by carina laterally. Gena glabrous and sparsely foveate, without medial vertical carina. Lower face foveate-reticulate. Anterior tentorial pits distinct. Clypeo-pleurostomal sulcus and epistomal sulcus form a smoothly curved arch. Clypeus transversely rugose and foveate posteriorly, and glabrous and finely punctate anteriorly, anterior third separated by a transverse inflection. Lateral occipital carina, reaching vertex. Occiput glabrous.

Anterior flange of pronotum longitudinally striate and punctate; submedian pronotal depressions distinctly separated medially. Anterior plate of pronotum foveate and densely punctate with pubescence medially, and glabrous laterally. Pronotum dorsomedially raised, but lower than mesoscutum; pronotal crest broadly emarginate medially; two submedian processes present immediately before crest. Lateral pronotal carina distinct, reaching pronotal crest dorsomedially. Lateral surface of pronotum transversely costate with superficial foveae set in rows between costae, and glabrate lateroposteriorly and lateroventrally. Dorsal pronotal area glabrous, present only along anterior half of posterior margin of pronotum. Mesoscutum slightly arched dorsally, transversely costate with superficial foveae set in rows. Scutellar sulcus divided by four longitudinal, equally strong carinae; mesoscutellum foveate-reticulate and raised posteriorly; posterior margin with a deep, wide triangular emargination in dorsal view and with conspicuous white pubescence. Axillar area with conspicuous white pubescence. Mesopleural triangle well defined ventrally by a smoothly curved carina and with conspicuous white pubescence. Median mesopleural impression percurrent, without transverse costae; both upper and lower mesopleuron glabrous; lower mesopleuron bordered by a distinct, straight carina ventrally. Metepisternum longitudinally costate and devoid of pubescence in upper part, conspicuously pubescent ventrally. Propodeum areolate-reticulate; lateral propodeal carina percurrent, distinctly curved posteriorly, and flattened inward dorsally to form a glabrous dorsal surface; median propodeal area glabrate with fine pubescence; median longitudinal carina distinct present anteriorly and absent posteriorly; two sinuated submedian longitudinal carinae present in posterior part of median propodeal area; transverse carina weak. Rs+M of forewing arising from anterior third of basal vein. Marginal cell 3.4 times as long as wide and 1.5 times as long as submarginal cell. Bulla on Sc+R 1 absent.

Abdominal petiole 0.57 times as long as wide in lateral view. Relative length of T3–8: 2.4:1.0:1.3:5.3:0.6:1.6; T3 glabrous with a row of setigerous punctures; T4–5 finely punctate; T6 mostly more or less densely, finely punctate, except with a median patch of long pubescence in larger punctures dorsolaterally; T7 finely punctate with a row of pubescence in larger punctures anteriorly; and T8 coarsely punctate-foveate with pubescence. Legs more or less densely pubescent. Apical teeth of metatibia short, stout, and blunt apically. 1mt/2–5mt 5 0.68.

MALE: Unknown.

Paramblynotus costaricanus is most similar to P. zonatus . They can be separated from P. braziliensis by having (1) vertex glabrous with sparse punctures; (2) submedian processes before pronotal crest not as prominent; (3) lateral sides of posterior part of mesoscutel- lum not abruptly narrowed behind dorsal lateral process; (4) forewing without a broad smoky band along outer margin; (5) T6 of female metasoma with tip distinctly not reaching as far as T7 posteriorly; dorsal margin abruptly curved posteriorly, and posterior margin slightly concave. P. costaricanus can be furhter separated from P. zonatus by having (1) lateral sides of pronotum largely glabrous and gena entirely glabrous, and (2) mesoscutellum posteriorly with a broad triangular emargination.

TYPE MATERIAL: HOLOTYPE: ♀, Costa Rica: Guanacaste: Santa Ross National Park, 1985-IV-17–27, collected by D. Janzen and I. Gauld (with Malaise trap), deposited in the Natural History Museum, London ( NHM). PARATYPES: 3♀♀. 2♀♀, locality and depository same as holotype, 1985-IV-17–27, collected by I. Gauld, and 1986-V-4–24, by D. Janzen and I. Gauld. 1♀, Guatemala: Zacapa (2 km from N. Santa Cruz), 1986-XI, collected by M. Sharkey (in dry river bed with Malaise trap), deposited in Canadian National Collection of Insects, Ottawa, Canada ( CNCI).

DISTRIBUTION: Costa Rica; Guatemala.

ETYMOLOGY: Named after holotype locality.