Paramblynotus obscurus, Liu, Ronquist & Nordlander, 2007

Liu, Zhiwei, Ronquist, Fredrik & Nordlander, Göran, 2007, The Cynipoid Genus Paramblynotus: Revision, Phylogeny, And Historical Biogeography (Hymenoptera: Liopteridae), Bulletin of the American Museum of Natural History 2007 (304), pp. 1-151 : 126-127

Paramblynotus obscurus , new species

FEMALE: Length 3.5–4.5 mm. Body and antenna entirely black. Legs black except tibiae and tarsi of fore- and middle legs yellow brown. Wings transparent. 1mt/2–5mt 5 0.60.

Antenna 13-segmented, flagellum filiform, not widened apically. Vertex foveate-reticulate. Eye extended laterally to as wide as outer margin of gena. Ocellar plate raised and foveate-reticulate and defined laterally by carina lined along interior side with a row of uniform large foveae; a glabrous triangular area present beneath anterior ocellus. Medi- an frontal carina briefly present between antennal sockets. Upper face rugose-foveate/punctate laterally; antennal scrobe distinctly depressed, glabrate and punctate with long, appressed pubscence, and defined by a carina laterally. Gena glabrate and foveatepunctate with dense, appressed pubescence; lower face and clypeus foveate-reticulate and densely punctate with dense appressed pubescence; anterior tentorial pits small and distinct; clypeo-pleurostomal sulcus and epistomal sulcus form a superficial, smoothly curved arch. Lateral occipital carina not reaching vertex. Occiput longitudinally carinate medially and glabrous laterally.

Anterior flange of pronotum finely coriarious; submedian pronotal depressions separated from each other medially; anterior pronotal plate glabrate and finely densely punctate with appressed pubescence. Pronotum raised dorsomedially; pronotal crest gradually raised medially into a peak higher than mesoscutum. Lateral pronotal carina distinct, present only along lower third of posterior margin of anterior pronotal plate. Lateral surface of pronotum foveate-reticulate and densely punctate with transverse carinate component and conspicuous ap- pressed pubescence. Dorsal pronotal area finely transversely striate, present along anterior fourth of posterior margin of pronotum. Mesoscutum flat dorsally and strongly transversely costate with superficial foveae set in rows and evenly distributed appressed pubescence. Scutellar sulcus divided by a single median longitudinal carina. Axillar area with conspicuous pubescence. Mesoscutellum foveate-reticulate; mesoscutellum broadly rounded posteriorly in dorsal view; lateral dorsal process visible, but indistinct in dorsal view. Mesopleural triangle conspicuously pubescent and well defined ventrally by smoothly curved carina. Median mesopleural impression percurrent and glabrous. Upper mesopleuron glabrous and with sparse, short pubescence anteriorly; lower mesopleuron glabrous and densely pubescent in ventral impression. Metepisternum areolate-reticulate in upper part, conspicuously pubescent ventrally, and with a large central elevated glabrous area. Lateral propodeal carinae percurrent, distinctly curved laterad and distinctly raised dorsally to form a broadly rounded lobe; median propodeal area glabrate; median longitudinal carina percurrent crossed by median transverse carina. Rs+M of forewing nebulous, arising from middle of basal vein. Marginal cell 2.2 times as long as wide and 1.5 times as long as submarginal cell. Bulla on Sc+R 1 absent.

Petiole 0.50 times as long as wide in lateral view. Tergum 8 completely covered by T7; relative size of abdominal T3–7: 1.9:1.0:1.2:2.7:0.8; T3–4 glabrous; T5 densely finely punctate; T6 finely and densely punctate with a narrow band of sparse pubescence across anterior two-fifths of the tergite; and T7 densely punctate, with a band of relatively dense pubescence dorsolaterally in anterior half and glabrous posteriorly. Apical teeth of metatibia slender and pointed apically. Apical protuberance of first metatarsomere reaching slightly beyond middle of second metatarsomere.

MALE: Body length 3.6 mm. Antenna 14-segmented. Terga 3–8 subequal with T3 and T5 larger than the others.

Paramblynotus obscurus differs from all other species of the genus except P. miniatus in having metatibia dorsally with dents. It differs from the latter in (1) forewing with

a broad dark gray band along outer margin; (2) upper metepisternum without a large, glabrous elevated median area; (3) head, pronotum, and metasomal T6–7 covered with conspicuous pubescence; and (4) body size much larger.

TYPE MATERIAL: HOLOTYPE: ♀, Borneo: North Borneo (Forest Camp, 19 km north of Kalabakan ), 1962-XI-12, Y. Hiroshima coll. ( BPBM). PARATYPES: 2♀♀, 1Oi. 1 Oi , Philippines: Palawan, Brooke’s Pt., Macagua (75 m), 1962-IV-1–4, M. Thompson coll. ( BPBM) ; 1♀, Malaysia: Sabah, Danum Valley (forest), 1986-X-23–XI-10, P. Eggleton coll. ( BMNH) ; 1♀, Sarawak, Gunung Buda , 64 km S Limbang (4 ° 13 9 N, 114 ° 56 9 E), 1996- XI-22–28, S.L. Heydon and S. Fung coll. ( UCDC) .

ADDITIONAL MATERIAL EXAMINED: 1Oi, Philippines: Mindanao (paratype of P. borneanus ) ( USNM).

DISTRIBUTION: Malaysia: Borneo; Indonesia: Sarawak; Philippines: Palawan.

ETYMOLOGY: From Latin, obscurus , meaning dark. The name describes the entirely dark color of its body.