Paramblynotus grossus, Liu, Ronquist & Nordlander, 2007

Liu, Zhiwei, Ronquist, Fredrik & Nordlander, Göran, 2007, The Cynipoid Genus Paramblynotus: Revision, Phylogeny, And Historical Biogeography (Hymenoptera: Liopteridae), Bulletin of the American Museum of Natural History 2007 (304), pp. 1-151 : 128-129

Paramblynotus grossus , new species

FEMALE: Length 6.0–8.0 mm. Body, antenna, and legs entirely black. Wings transparent; forewing with a rectangular macula along the middle of anterior margin, covering marginal cell, distal third of first submarginal cell, and basal part of second submarginal cell anteriorly. 1mt/2–5mt 5 0.

Antenna 13-segmented, flagellum filiform, not widened apically. Vertex foveate-reticulate. Eye not extended laterally, outer margin of gena expanded much beyond outer margin of eyes. Ocellar plate distinctly raised and foveate and not defined laterally by carina. Median frontal carina briefly but strongly present between antennal sockets. Upper face rugose-foveate/punctate laterally; antennal scrobe distinctly depressed and coarsely punctate with distinct pubescence, longitudinally carinate posteriorly, and distinctly defined by carina laterally. Gena glabrate, and foveate and punctate with sparse pubescence; lower face and clypeus foveate and densely punctate with apparent transverse costate component and appressed pubescence; ante- rior tentorial pits distinct; clypeo-pleurostomal sulcus and epistomal sulcus form a superficial trapezoid. Lateral occipital carina reaching to middle height of gena. Occiput longitudinally carinate medially and glabrous laterally.

Anterior flange of pronotum glabrous; submedian pronotal depressions separated medially; anterior pronotal plate glabrous medially and otherwise finely densely punctate with appressed pubescence (especially so laterally). Pronotum raised dorsomedially; pronotal crest gradually raised medially into a peak higher than mesoscutum. Lateral pronotal carinae distinct, present in lower two-thirds of posterior margin of anterior pronotal plate. Lateral surface of pronotum foveate-reticulate with transversely costate components, and distinctly pubescent. Dorsal pronotal area glabrate, present along anterior third of posterior margin of pronotum. Mesoscutum almost flat dorsally and transversely costate with superficial foveae set in rows and with sparse, evenly distributed appressed pubescence. Scutellar sulcus divided only by a median longitudinal carina. Pubescence of axillar area distinct but not conspicuous. Mesoscutellum foveate-reticulate; mesoscutellum broadly rounded posteriorly in dorsal view; lateral dorsal process not apparent in dorsal view. Mesopleural triangle conspicuously pubescent and well defined ventrally by smoothly curved carina. Median mesopleural impression percurrent and glabrous. Upper mesopleuron glabrous; lower mesopleuron also with dense pubescence in ventral impression. Metepisternum areolatereticulate to rugose in upper part, conspicuously pubescent ventrally, and with an elevated, glabrous median area. Lateral propodeal carinae percurrent, nearly parallel to each other, and strongly raised dorsally to form in middle a distinct triangular process; median propodeal area areolate with sparse pubescence; median longitudinal and transverse carinae hardly distinguishable from ridges of areolate structure. Rs+M of forewing nebulous in basal third, arising from middle of basal vein. Marginal cell 3.1 times as long as wide and 1.1 times as long as submarginal cell. Bulla on Sc+R 1 absent.

Petiole 0.45 times as long as wide in lateral view. Tergum 8 completely covered by T7; relative size of abdominal T3–7: 2.0:1.0:2.0.:2.4:1.5; T3–4 glabrous; T5 glabrous and finely punctate; T6 finely and densely punctate dorsolaterally and with a narrow band of sparse pubescence in the middle; T7 punctate with pubescence dorsolaterally in anterior third. Apical teeth of metatibia slender and pointed apically. Metatibia with two to three dorsal (posterior) dents. Apical process of first metatarsomeres reaching to middle of second metatarsomere.

MALE: Unknown.

Paramblynotus grossus differs from all other species of the punctulatus group except P. venoforticulus , n.sp. by the presence of a rectangular macula along the middle of anterior margin of forewing. It differs from P. venoforticulus by its conspicuously expanded gena behind eye. The combination of robust shape, big size, and entirely black body color also make the identification of the species easy.

TYPE MATERIAL: HOLOTYPE: ♀, Papua New Guinea: Bulolo, Mankila , 1981-XII-24, H. Roberts coll. ( NHM). PARATYPES: 15♀♀. 5♀♀, data as holotype ( NHM) ; 1♀, New Guinea, Wau (1,200 m), 1955-XII-22, I. and M. Sedlacek coll. ( NHM) ; 1♀, Laos: Vientiane Prov., Ban Van Eue, 1965-VII-31, Native Collector coll. ( BPBM) ; 2♀♀, Malaysia: Sabah, Sipitang: Mendolong , 1989-III-8– 14, S. Adebratt, ( ZMLU) ; 6♀♀, Indonesia: Sulawesi: Kunogon-Bone, 1984-IV–V, Martin coll. ( NHM), 1985-XI-27–XII-4 ( NHM), Dumoga-Bone (200 m), 1985-XI-16–23, C. v. Achterberg coll. ( NNMN) ; Sumatra, Aceh, G. Leuser National Park, Ketambe Research Station , 1989-XI – 1990-II, C.D. Darling coll. ( ROM) .

DISTRIBUTION: Indonesia: Sulawesi, Sumatra; Malaysia: Borneo; Laos: Vientiane; Papua New Guinea.

ETYMOLOGY: From Latin, grossus , big, coarse, referring to the big size and robust shape of the species.

BIOLOGY: Specimens from Papua New Guinea, including the holotype, were collect- ed on Syzigium sp. log.