Promalactis ramivalvata Wang, 2020

Wang, Shuxia & Liu, Chen, 2020, Taxonomic study of the genus Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) IX. The maculosa species-group, with descriptions of eighteen new species, Zootaxa 4890 (1), pp. 38-66 : 50-52

Promalactis ramivalvata Wang , sp. nov.

( Figs 13 , 30 , 44 )

Type material. CHINA, Yunnan: Holotype ♂, Mt. Weibao (25.18°N, 100.32°E), Weishan, 2200 m, 20.VII.2001, leg. HH Li & XP Wang, slide No. W01226 . Paratypes (34♂, 28♀): 10♂, 3♀, same data as holotype, slide Nos. DZH10183 ♀, DZH10184 ♂, DZH10186 ♂, W01238 ♀; 6♂, 11♀, Mt. Weibao , Weibao County, 2244 m, 21– 24.VII.2013, leg. SR Liu et al., slide Nos. HXM13170 ♂, HXM13193 ♂, JYY17720 ♀; 1♀, Mt. Xiaohei, Longling , 1974 m, 19.VII.2013, leg. SR Liu et al., slide No. LC19477 ; 1♂, Yexiang Village, Xishuangbanna , 762 m, 19.VII.2014, leg. KJ Teng et al .; 17♂, 23♀, Mt. Weibao, Dali , 2205 m, 30.VII–3.VIII.2014, leg. KJ Teng et al., slide Nos. HXM14146 ♂, JYY17196 ♂, JYY17197 ♂, LC19430 ♀, LC19435 ♀, LC19438 ♀.

Diagnosis. The new species is similar to P. quadriloba Du et Wang, 2013 , but it can be distinguished in the male genitalia by the lingulate gnathos, the sub-ovoid juxta without lateral arms, and the valva lacking a spine-like ventrodistal lobe. In P. quadriloba , the gnathos is rectangular, the juxta has lateral arms, and the valva has a ventrodistal lobe ( Du & Wang 2013: 31, fig. 21).

Description. Adult ( Fig. 13 ). Wingspan 7.5–12.0 mm. Head with vertex silvery white, frons dark brown, oc-ciput dark brown mottled with silvery white. Labial palpus with basal segment grey; second segment grey in basal 2/3, black in distal 1/3; third segment black except basal 1/3 and apex white, slightly shorter than second segment. Antenna with scape white, pecten dark brown; flagellum black alternated with white on dorsal surface, dark brown on ventral surface. Thorax and tegula rust brown. Forewing yellowish brown; markings white edged with black scales: costal spot large, quadrate, placed beyond middle, reaching posterior angle of cell, its outer margin sinuate; cell with a wide stripe extending from basal 1/4 obliquely downward to basal 2/5 of fold; fold with a dot at base; dorsum with three short streaks: first streak from basal 1/5 extending to white dot at base of fold, second streak from basal 2/5 to basal 2/5 of fold, third streak extremely slender, from basal 3/5 oblique outward to posterior angle of cell; apical spot triangular, with dense black scales on inner margin; tornal spot grey, diffused to posterior angle of cell, tinged with a few white scales; cilia deep ocherous yellow on basal half, ocherous yellow on distal half, mottled with black scales around tornus. Hindwing and cilia grey. Foreleg black, tibia with a small white spot at base and middle dorsally, with a tuft of white scales apically, tarsus white at apices of basal two tarsomeres; midleg grey ventrally, black dorsally, tibia with a white spot at base dorsally, with a tuft of white scales basally and apically, tarsus white at apices of basal two tarsomeres dorsally; hindleg yellow, tibia grey dorsally, tarsus black dorsally except white at apices of basal two and fourth tarsomeres and on entire apical tarsomere.

Male genitalia ( Fig. 30 ). Uncus with basal half sub-rounded, distal half uniformly narrow, rounded apically.

Gnathos shorter than uncus; mesial plate broad lingulate, rounded apically; basal arm band-like, slightly shorter than mesial plate. Tegumen divided from posterior 1/4; lateral arm distinctly narrowed to rounded apex. Valva narrow at base, gradually widened to apex, with three apical lobes: dorsal lobe greatly arched ventrad, exceeding ventral lobe apically, its basal half smoothly uniform, distal half with dense setae and short spines, narrowed to pointed apex, like ear of wheat, middle lobe from near base of dorsal lobe, clavate, dilated and with dense setae distally, extending almost straightly outward, close to and shorter than dorsal lobe, ventral lobe as long as middle lobe, wide basally, narrowed to rounded apex distally, setose distally, serrate ventrobasally; costa narrow and short; sacculus wide, uniformly wide, oblique and serrate apically. Saccus short, about 3/5 length of uncus, rounded apically. Juxta broad and short, produced in large tongue-shaped lobe posteriorly, widened and triangularly produced outward lateromedially. Aedeagus slightly shorter than valva; cornutus spine-like, about 1/5 length of aedeagus, located distally.

Female genitalia ( Fig. 44 ). Apophyses anteriores about half length of apophyses posteriores. Eighth sternum very large, with dense setae, posterior margin deeply concave, forming two sub-triangular lateral processes. Ostium bursae small, deeply incised posteromedially. Antrum funnel-shaped, more sclerotized laterally. Ductus bursae membranous, as long as corpus bursae, posteriorly expand and with two claw-like plates, each plate with strong spines arising from a short stem. Corpus bursae large, elliptical; signum absent.

Distribution. China (Yunnan).

Etymology. The specific epithet of this new species is derived from the Latin ramus and valva, referring to the branched apex of the valva in the male genitalia.

Figures

FIGURES 13‒18. Adults of Promalactis spp. 13, P. ramivalvata, holotype, ♂; 14, P. aculeiformis, holotype, ♂; 15, P. apicitriangula, paratype, ♂; 16, P. curvispinosa, paratype, ♂; 17, P. medimacularis, holotype, ♂; 18, P. serraticostalis, holotype, ♂. Scales = 2 mm, except 15 scale = 1 mm.

FIGURES 25‒30. Male genitalia of Promalactis spp. 25, P. plicata, holotype, slide No. JYY17788; 26, P. bomiensis, holotype, slide No. LC19226, 26a, enlarged cornutus; 27, P. brevipalpa, holotype, slide No. DZH12288, 27a, enlarged cornutus; 28, P. libona, holotype, slide No. JYY17787, 28a, enlarged cornutus; 29, P. medilargissima, holotype, slide No. LC19307, 29a, enlarged cornutus; 30, P. ramivalvata, paratype, slide No. DZH10184. Scales = 0.2 mm.

FIGURES 42‒48. Female genitalia of Promalactis spp. 42, P. libona, paratype, slide No. JYY17722, 42a, enlarged signa, slide No. JYY17723; 43, P. medilargissima, paratype, slide No. LC19429; 44, P. ramivalvata, paratype, slide No. LC19438, 44a, enlarged ductus bursae; 45, P. curvispinosa, paratype, slide No. DZH11037; 46, P. medimacularis, paratype, slide No. JYY17829; 47, P. serraticostalis, paratype, slide No. DZH12114; 48, P. tricuspidata Wang et Li, 2004, slide No. JYY17977. Scales = 0.2 mm.