Triclistus amazopeikkus, Sääksjärvi, Ilari Eerikki & Bordera, Santiago, 2015

Sääksjärvi, Ilari Eerikki & Bordera, Santiago, 2015, A bizarre new species of Triclistus Förster (Hymenoptera: Ichneumonidae; Metopiinae) from Amazonia, Zootaxa 3955 (4), pp. 595-600 : 596-600

publication ID

https://doi.org/ 10.11646/zootaxa.3955.4.10

publication LSID

lsid:zoobank.org:pub:B482EBE3-A537-46AF-AE8D-088C1936D565

DOI

https://doi.org/10.5281/zenodo.6118509

persistent identifier

https://treatment.plazi.org/id/039CB01D-FFA0-FF8B-FF72-FEB09BC563E3

treatment provided by

Plazi

scientific name

Triclistus amazopeikkus
status

sp. nov.

Triclistus amazopeikkus sp. nov.

( Figs 1 View FIGURE 1 A–B, 2A–F, 3A–E)

Diagnosis. The species is easily identified as Triclistus by the specialized form of the subgenital plate which is modified to support the thin ovipositor ( Fig. 3 View FIGURE 3 E). In addition, the species is dark colored ( Figs 1 View FIGURE 1 A–B) whereas most Colpotrochia Holmgren / Cubus Townes are mostly orange and black in coloration. Triclistus amazopeikkus sp. nov. can be easily distinguished from all other species of the genus by its extremely specialised face which is centrally projecting and forming an extraordinary sub-pyramidal prominence, ending anteriorly with two apical points ( Figs 1 View FIGURE 1 B, 2B–C). These in turn arise from lamellae, which are separated by vertical grooves from the antennal sockets ( Fig. 2 View FIGURE 2 B).

Description. Female: Body size, 4.9–5.4 mm. Fore wing length 3.2–3.6 mm.

Head: antennae with 20 flagellomeres, gradually narrowed towards the apex ( Figs 1 View FIGURE 1 A, 1B), first, second and subapical flagellomeres 5.0–5.3, 1.4 and 2.0 × as long as centrally broad, respectively; combined face and clypeus 1.0–1.1 × as long (from clypeal apex to level of antennal sockets) as wide (shortest distance between eyes) ( Fig. 2 View FIGURE 2 D); face centrally projecting, forming an extraordinary sub-pyramidal prominence ( Fig. 2 View FIGURE 2 A–C), ending apically with two points which arise from lamella and both vertical grooves separating it from antennal sockets ( Fig. 2 View FIGURE 2 B), shiny very densely and coarsely punctate at the prominence, gradually sparse ventrally ( Fig. 2 View FIGURE 2 C); prominence with strong longitudinal striae ventrally ( Fig. 2 View FIGURE 2 E); clypeus smooth and shiny, clypeal margin concave ( Fig. 2 View FIGURE 2 C); labrum exposed when mandibles closed ( Fig. 2 View FIGURE 2 C), 2.3–2.5 × as wide as long, shiny, with few sparse setiferous puctures, apical rim somewhat bilobed; mandible strongly tapered, somewhat bent upward at the apex, with upper tooth conspicuously longer than lower tooth; malar space 1.0–1.2 × as long as basal mandibular width; lateral ocellus separated from compound eye by 1.5–1.7 × maximum ocellar diameter; distance between posterior ocelli about 1.0 × maximum ocellar diameter ( Fig. 2 View FIGURE 2 B); head in dorsal view with gena slightly rounded and moderately constricted behind eyes ( Fig. 2 View FIGURE 2 B); gena, on lateral view 0.7–0.8 × as long as compound eyes, upper part mostly smooth and shiny, sparsely punctate behind ocelli, lateral part smooth and shiny, with a line of 5–6 brown small sensillae close and parallel to the posterior margin of eye, posteriorly with a group of 6–7 sensillae ( Fig. 2 View FIGURE 2 F, arrows); lower part gradually densely punctate ( Fig. 2 View FIGURE 2 C); vertex quite prominent, coarsely and densely punctate in interocellar space ( Fig 2 View FIGURE 2 A–B); frons coarsely and densely punctate, very leaning forward and confluent with the dorsal part of the facial subpiramidal prominence, crest between antennal toruli wide, laterally with two deep grooves to house the antennae when directed backwards ( Fig. 2 View FIGURE 2 B).

Mesosoma : pronotum shiny, very densely punctate on upper medial and lateral part, setae long; femoral concavities smooth and hairless; mesoscutum flat, coarsely and densely punctate ( Fig. 1 View FIGURE 1 A); notauli extending beyond centre, very weak ( Fig. 1 View FIGURE 1 B); scuto-scutellar groove deep and smooth; scutellum slightly convex, almost flat, with longitudinal lateral carinae reaching to about 0.4 × its length ( Fig. 1 View FIGURE 1 B); mesopleurum smooth and shiny, densely punctate anteriorly, impunctate posteriorly ( Figs 1 View FIGURE 1 A, 3A); mesosternal region with a flattened subtriangular process projecting backwards over the basis of the mid coxa ( Fig. 3 View FIGURE 3 A); metapleuron smooth and shiny, with isolated setae in anterior and posterior area, posterior upper corner forming a right angle ( Fig. 3 View FIGURE 3 A); submetapleural carina smooth, anteriorly expanded into a conspicuous triangular lobe ( Fig. 3 View FIGURE 3 A). Propodeum quite long and slightly convex, weakly declivitous and abruptly down-turned posteriorly with longitudinal carinae strong; confluent area basalis and area superomedia smooth and shiny, hairless ( Fig. 3 View FIGURE 3 B), confluent area externa and dentipara smooth and shiny with fine and sparse setiferous punctures ( Fig. 3 View FIGURE 3 B), confluent area spiracularis and area lateralis slightly rugulose-granulate, with very dense white pubescence ( Fig. 3 View FIGURE 3 A); posterior transverse carina strong; area peciolaris and area posteroexterna confluent, slightly rugulose-granulate with white pubescence; area coxalis forming a deep concavity with white setae ( Fig. 3 View FIGURE 3 A); spiracle elongate ( Fig. 3 View FIGURE 3 A). Front leg with tibial spur stout and claw-like ( Fig. 3 View FIGURE 3 C, arrow); mid leg with anterior tibial spur 0.8–0.9 x length posterior spur, not specialized; hind leg with anterior spur short and stout about 0.5 × length of posterior spur, both spurs strongly sclerotized apically, claw-like ( Fig. 3 View FIGURE 3 D); distal hind tarsomere without apparent hooked lobe on the inner side. Fore wing with vein 3 rs-m absent.

Metasoma: tergite I smooth and shiny, with fine and very sparse setiferous puctures laterally, 1.45 × as long as posteriorly broad, lateral longitudinal carinae reaching to spiracle, lateromedian carinae extending 0.5 × length of tergite ( Fig. 3 View FIGURE 3 B); tergite II smooth and shiny, with very fine and moderately dense setiferous punctures, denser laterally, about 0.92 × as long as posteriorly broad; rest of tergites with same microsculpture but punctures somewhat denser in last ones; subgenital plate with hind margin modified into deeply V-shaped concavity centrally ( Fig. 3 View FIGURE 3 E).

Colouration ( Figs 1 View FIGURE 1 A–B). Black. Antennae, central part of mandibles front and mid legs, and hind tibia basally yellow; hind coxa and femur, and distal apex of front and hind tibial spurs, dark brown; hind trochanters, distal part of hind tibia, hind tarsi, labrum and base of mandibles light brown; maxillae and labium entirely white.

Male. Unknown

Etymology. We are pleased to name the species as “ amazopeikkus ” which is a combination of names suggested by the children studying in the Kuparivuori School (class 4B), Naantali, Finland. The word “ amazo ” refers to Amazonia and the word “ peikkus ” is derived from the Finnish word “Peikko” (Peikko is a troll in Finnish). The name “ amazopeikkus ” means “the troll of Amazonia”. In our opinion, it describes well the extraordinary morphology and unknown habits of this species.

Material examined. Holotype female: Peru, Department of Loreto, National Reserve of Allpahuayo- Mishana, Malaise trap E1 (12), 14.IX.2000, Ilari E. Sääksjärvi et al. leg. ( UNSM).

Paratype female: Peru, Department of Loreto, National Reserve of Allpahuayo-Mishana, Malaise trap C1, VIII–XII.1998, Ilari E. Sääksjärvi et al. leg. ( ZMUT).

UNSM

University of Nebraska State Museum

ZMUT

University of Tokyo, Department of Zoology

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Hymenoptera

Family

Ichneumonidae

Genus

Triclistus

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