Najash, APESTEGUIA AND ZAHER, 2006

Zaher, Hussam, Apesteguía, Sebastián & Scanferla, Carlos Agustín, 2009, The anatomy of the upper cretaceous snake Najash rionegrina Apesteguía & Zaher, 2006, and the evolution of limblessness in snakes, Zoological Journal of the Linnean Society 156, pp. 801-826 : 803

NAJASH APESTEGUÍA AND ZAHER, 2006

Emended diagnosis: A snake nearly 2 m long with robust hindlimbs and a sacrum, tip of dentaries with a medially projected facet that bears a straight anteroposteriorly directed margin, suggesting a tightly connected mandibular symphysis, lack of dentary shelf, prootic exposed dorsally between the otooccipital and parietal, lack of laterosphenoid, developed laterally projected basipterygoid process, lack of a crista circumfenestralis, robust stapedial footplate, single large parazygantral foramen on vertebrae, arqual ridges on middle and posterior presacral vertebrae, and blunt haemapohyses on caudal vertebrae. It exhibits the following autapomorphies: (1) a thick splenial; (2) strongly concave ventral surface of the parasphenoid rostrum, forming a deep and straight gutter; (3) strongly faceted condition of the neural arch laminae; (4) enlarged and blade-like femoral trochanter.