Lomographa bimaculata (Fabricius, 1775)

Beljaev, E. A., 2020, New contribution into concept of the tribe Deveniliini (Lepidoptera: Geometridae, Ennominae) based on skeleton-muscular anatomy of the male genitalia, Far Eastern Entomologist 420, pp. 1-13 : 7-9

Lomographa bimaculata (Fabricius, 1775)

Figs 7–9

MATERIAL. Russia: Primorskii krai, 20km SE Ussuriisk, Gornotayozhnoe,

43°42′ N, 132°09′ E, 1.VI 1989, 3♂, E.A. Beljaev leg.

1775): 7 – the male genitalia, ventroposterior view, muscles m3(2), m4 and m6(5) are not shown on the right, and all paired muscles are not shown on the left; 8 – area of tegumen,

vinculum and valva articulation, ventroposterior view, muscles m5(7) are not shown; 9 –

phallus, lateral view, muscles m6(5) and m7(6) are not shown on the right; 10 – Cabera pusaria (Linnaeus, 1758) , area of tegumen, vinculum and valva articulation, medial view,

muscles m1, m6(5), m7(6), m8(3) are not shown. Abbreviations: see Fig. 1 .

ded on tegumen and vinculum by thin narrowing. Lateral lobes of tegumen are narrow, ribbon-like, dorsally connected by short bar-like bridge. Vinculum is thin,

saccus is not developed. Uncus is thin, long, falcate, socii are large, membranous.

Gnathos is weak, in form of thin semiring. Valvae are long and wide, lobe-like,

with broad weakly defined cucullus. Costa is moderately sclerotized, wide, in basal half with long longitudinal opening dorsally. Hemitranstilla in form of bar-like lobe with almost parallel sides; left and right hemitranstillae are touching medially.

Cuiller (sensu Beljaev, 2008: by origin it is a distal sclerite of sacculus, which in many geometrids remains as a more or less broad plate in the ventrobasal portion of valvula) elongate, large, occupies most of medial side of valvula, distally finished with lobe-like free distal process. Clasper is integrated with cuiller and do not expressed morphologically. Sacculus is long, flat, moderately sclerotized. Juxta is large, broad and long, dorsally with two large lobe-like protuberances, cristae integrated in lateral sides of juxta. Labides are lacking or integrated with juxta in form of two small digital processes located just laterad of the dorsal protuberances.

Aedeagus is relatively small, cylindrical, with prominence caecum, vesica bears brush-like group of needle cornuti protruded from the aedeagus.

The musculature. Depressors of the uncus, m1, are moderate, extending from ventral corners of tegumen, distally attached to the ventral ribs of lateral sides of uncus just before the base of distal process of uncus. Anal cone retractors, m2(10),

are extending from anterior portion of lateral lobes of tegumen. Dorsal abductors of valvae, m3(2), are short, extending from the anterior rib of tegumen near its ventral corner to ventral margin of distal half of hemitranstilla. Adductors of valvae, m4,

are also short, extending from small dilation of dorsal corners of vinculum to the lateral side of basal half of the hemitranstilla. Flexors of valvae, m5(7), are short,

moderately wide, proximally attached partly to the medial wall of the sacculus,

partly to the lateral edge of juxta, distally attached to ventral half of proximal rib of cuiller. Ventral extensor of valva, m8(3), is unpaired, wide, extending from the most ventral portion of vinculum to near the 1/3 length of juxta. Protractors of the phallus, m6(5), are thin, extending from near the dorsal corners of vinculum just ventrad of m4. Retractor of the phallus, m7(6), unpaired, wide, extending from ventroanterior rib of vinculum to near the 1/3 length of ventral side of aedeagus.

Retractor of vesica, m21, is present, but not shown.

Figures

Figs 7–10. Musculature of the male genitala. 7–9 – Lomographa bimaculata (Fabricius,

Figs 1, 2. Devenilia corearia (Leech, 1891), musculature of the male genitala. 1 – the