Synegia ichinosawana (Matsumura, 1925)

Beljaev, E. A., 2020, New contribution into concept of the tribe Deveniliini (Lepidoptera: Geometridae, Ennominae) based on skeleton-muscular anatomy of the male genitalia, Far Eastern Entomologist 420, pp. 1-13 : 6-7

Synegia ichinosawana (Matsumura, 1925)

Figs 5, 6

MATERIAL. Russia: Sakhalin, Yasnomorskoe vicinity, N slope of Sokhonda mountain , 46°44'52"N, 141°55'01" E, 70 m asl, 21–23.VII 2019, 2♂, E.A. Beljaev leg.

MALE GENITALIA ( Figs 5, 6 ). The skeleton. Genital segment is clearly divided on tegumen and vinculum by thin narrowing. Lateral lobes of tegumen are narrow,

ribbon-like, dorsally distanced and connected by extended bar-like bridge. Vinculum is very wide, strongly sclerotized, with complicate internal system of stiffening.

Uncus is large, conic, but moderately sclerotized, socii membranous. Gnathos is weak, in form of thin semiring with a pair of lobe-like dilations medioventrally.

Valvae are long, narrow, dorsally connected with vinculum by strong articulation formed by posterior dilation of vinculum and anterior thickening of lateral wall of the costa valvae. Costa is very wide, well sclerotized, finished by pointed spine.

Hemitranstilla in form of firm and long digital process, left and right hemitranstillae are touching medially. Ventrad of hemitranstilla, costa extends in broad lobe-like medial dilation. Cucullus broadly spreads along the costa almost to clasper, weakly delimited from valvula. Clasper is an arched narrow thickening of proximal edge of valvula. Sacculus is relatively short, well sclerotized, somewhat inflated. Juxta is simple, moderate in size, conic. Labides are lacking. Aedeagus is relatively small,

groove-like, membranous dorsally, vesica with a single strong straight subulate cornutus.

The musculature. Depressors of the uncus, m1, are moderate, extending from ventral corners of tegumen, distally attached to the anterior margin of the ventral wall of distal process of uncus. Anal cone retractors, m2(10), are extending from anterior rib of lateral lobes of tegumen remote from their dorsal connection. Dorsal abductors of valvae, m3(2), are strong and long, extending from the dorsal portion of anterior rib of lateral lobes of tegumen to membranous thickening ventad to distal half of hemitranstilla. Adductors of valvae, m4, are very strong, wide and long, extending ventrally from deep lateroanterior invaginations of vinculum to medial deletion of costa. Judging by the topology and attachment of the m4, this dilation should be a modified ventrobasal portion of hemitranstilla. Flexors of valvae, m5(7), are short, moderately wide, proximally attached to the basal corner of sacculus, distally extending to clasper. Ventral extensor of valva, m8(3), is unpaired, powerful, extending proximally from dorsal protrusion on saccus, distally attaching to near the top of juxta. Protractors and retractors of the phallus, m6(5)

and m7(6), are paired, thin, first ones extending from near the dorsal corners of vinculum and distally attached to lateral sides of anterior opening of aedeagus, and second ones extending from ventolateral sides of posterior rib of vinculum to the middle of ventral side of aedeagus. Retractor of vesica, m21, is present, but not shown.

NOTE. In the male genitalia skeleton, S. ichinosawana differs from Synegia botydaria Guenée, 1857 , the type species of the genus Synegia , by absence of a process near the base of costa, and presence single cornutus on vesica instead of two (see Holloway, 1994). But other structures conform well, so the male genitalia musculature of S. ichinosawana can presents its pattern in the genus.

Figures

Figs 3–6. Musculature of the male genitala. 3, 4 – Platycerota incertaria (Leech, 1891): 3 –