Calosoma depressicolle

Häckel, Martin, Heinz, Walter & Sehnal, Rostislav, 2011, Revision of the Mexican subgenus Carabomimus Kolbe, 1895 (Coleoptera: Carabidae: Calosoma Weber, 1801): C. depressicolle, C. flohri and C. digueti species groups, Zootaxa 2908, pp. 45-63 : 47-48

Key to the Calosoma depressicolle complex

1 Apex of aedeagus flag-shaped (characteristic of entire subgenus) and with transverse process conspicuous in vertical plane (Plates 3–4: Figs. 10, 13, 14–16, 18, 19. Plate 5: Fig 22 B, C; Fig 23 A, B; Fig 24); N Colima, S Jalisco, Michoacán................................................................................................. flohri group 2.

- Apex of aedeagus also flag-shaped but without transverse process in vertical view (Plates 1–2: Figs. 1, 2, 5, 7–9. Plates 5–6: Fig. 22 A, D; Fig. 23 C–E). Pronotum smooth or finely punctured at base and sides; Distrito Federal, México, Morelos........................................................................................ depressicolle group 6.

2 (1) Transverse process of aedeagus short and arcuate (Plate 4: Figs. 18, 19 ; Fig. 22 C; 23 B); NE Michoacán........ C. lesnei .

- Transverse process of aedeagus longer, trapezoidal (Plates 3–4: Figs. 10, 13, 14–16. Plates 5–6: Fig. 22 B; Fig. 23 A; Fig. 24); N Colima, S Jalisco, W Michoacán................................................... C. flohri ; four subspecies 3.

3 (2) Entire head and pronotum densely punctured; body smaller, 17.5– 21 mm (Plate 3: Fig. 10)................ C. flohri flohri .

- Head and pronotum smooth or coarsely punctured only near base.

4 (3) Elytral sculpture smooth or only coarsely punctured, without visible intervals, body smaller, 18–21 mm (Plate 3: Figs. 12–14 )....................................................................................... C. flohri gebieni .

- Elytral sculpture well expressed, with visible intervals, body larger, 20–24 mm.

5 (4) Head and pronotum near base coarsely punctured (Plate 4: Figs. 16, 17)........................... C. flohri zirostense .

- Head and pronotum near base almost smooth (Plate 4: Fig. 15)................................. C. flohri colimaense .

6 (1) Body smaller and wider (16–18 mm). Aedeagus with rounded termination of diameter so small that it appears pointed unless magnified (Plate 2: Figs. 7, 8 . Plates 5–6: Fig. 22 D; Fig. 23 E), and flag-shaped thickening weakly indicated, discernible only under magnification. Elytra with large, irregularly spaced punctures which interrupt neighboring, finely punctured intervals. Only C México: Volcán Nevado de Toluca.......................................................... C. altipeta .

- Body somewhat larger (length of smallest specimens exceeds 17 mm) and narrower; apex of aedeagus with flag-shaped thickening visible without magnification (Plates 1–2: Figs: 1–6, 9).................................................. 7.

7 (6) Apex of aedeagus in lateral view with pronounced step-like thickening; body larger, 20–24 mm (Plate 2: Fig. 9); E México: Río Frío; W Puebla............................................................................ C. sylvestre .

- Apex of aedeagus with weak thickening only, which viewed laterally narrows toward terminus gradually, without any step-like break (Plates 1–2: Figs. 1, 2, 5. Plates 5–6: Fig. 22 A; Fig. 23 C, D); body smaller, 16–20 mm; W, C and SE areas of México, E Michoacán............................................................ C. depressicolle , two subspecies ..... 8.

8 (7) Pronotum completely smooth, including sides and base. Elytral striae more regular, intervals flat or slightly convex, without scattered punctures or in some specimens with a few punctures only on sides (Plate 1: Figs. 1–4. Plate 6: Fig. 23 C); C México, Distrito Federal, Morelos........................................................ C. depressicolle depressicolle .

- Pronotum with a few punctures on sides and base. Elytral striae slightly disarrayed near apex (Plate 2: Figs. 5, 6. Plates 5–6: Fig 22 A; Fig 23D), deeper; intervals convex, with large, scattered punctures; foveae not always obvious; W México, E Michoacán............................................................................... C. depressicolle asper .

Figures

FIGURE 22. Aedeagus of Mexican Calosoma from central México. (A – D) Calosoma depressicollis complex (A, C. d. asper; B, C. d. flohri; C, C. d. lesnei; D, C. d. altipeta). E, C. digueti Lap. F, C. gebieni Breun. G, C. orizabae Jean. H, C. laevigatus Chaud. I, C. costipennis Chaud. J, C. striatulus Chevr. K, C. politus Chaud. L, C. diminutus Bat. M, C. diminutus morelianus Bat. (original illustrations sensu Jeannel 1940).

FIGURES 18 – 19. Calosoma (Carabomimus) lesnei Breuning, 1931. (18) Habitus of holotype, male. (19) Male from Volcán San Andres: Laguna Verde, E Michoacán. Bar equals 10 mm.

FIGURES 12 – 14. Calosoma (Carabomimus) flohri gebieni Breuning, 1931. (12) Female from env. of Pátzcuaro, W Michoacán. (13) male from 6.2 miles W Quiroga, W Michoacán. (14) Male from inbetween Cherán and Carapán (W Michoacán). Bar equals 10 mm.

FIGURES 7 – 8. Calosoma (Carabomimus) altipeta (Jeannel, 1940). (7) Habitus of lectotype male. (8) Male from Volcán Nevado de Toluca (central México).