Scorzonera undulata f. luteola Pau

Nualart, Neus, Soriano, Ignasi, Prieto, David Pérez & Ibáñez, Neus, 2021, Catalogue and typification of the Moroccan taxa published by Carlos Pau, Phytotaxa 519 (1), pp. 1-94 : 70-75

Scorzonera undulata f. luteola Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 355 (1924a)

= Scorzonera undulata Vahl

Ind. loc.:—“de Tánger a Tetuán”

Type:—not seen

1,2,3 Scrophularia mellifera var. vidalii Pau in Vidal, Bol. Soc. Esp. Hist. Nat. 25: 342 (1925)

= Scrophularia sambucifolia L. subsp. sambucifolia

Ind. loc.:—“Bu Allal”

Type:—not traced

Note:— APD (2020) cites this taxon as S. sambucifolia var. vidalii Pau , but this should be considered a combination published by Jahandiez & Maire (1934: 682) according to Art. 41.3 ( Turland et al. 2018).

2,3 Scrophularia trisecta Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 369 (1924a)

= Scrophularia sambucifolia L. subsp. sambucifolia

Ind. loc.:—“Rincón de Medik”

Lectotype (designated by Ortega Olivencia & Devesa 1996: 298):— Rincón de Medik , 7 May 1921, Pau s.n. (lectotype MA 111462 !; isolectotype BC 43750 !)

Notes:—The MA specimen was identified as Scrophularia sambucifolia subsp. sambucifolia by Ortega Olivencia in 1995.

1 Sedum album var. minutiflorum Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 327 (1924a)

= Sedum album L.

Ind. loc.:—“Beni Hozmar”

Holotype:—Beni Hozmar, May 1921, Pau s.n. ( MA 51284 ! [http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00051284])

Note:— The MA specimen was labelled as a spontaneous plant by Pau. In Herb. BC there is also a specimen from the same locality and date, but with the indication “cultivated” ( BC 23132 !) and so, not considered as type material. This cultivated gathering was distributed in the Sennen exsiccata Plantes d’Espagne 1934 nº 9181 (BCSennen 80823!, MA 51285 !). Although Byalt in 2005 labelled MA 51284 as isotype and MA 51285 as type, only the spontaneous plants should be considered original material .

1 Sedum cadevallianum Pau , in sched. ( MA 51854! [http://colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00051854]), nom. nud.

=?

Note:—This specimen bears a small sample without leaves, so determination has not been possible.

1,3 Senecio crassifolius var. latisectus Pau & Font Quer in Font Quer, Iter Marocc. 1929 nº 449 (1930a), in sched.

= Senecio leucanthemifolius subsp. mauritanicus (Pomel) Greuter

Ind. loc.:—“Hab. in umbrosis maritimis promontorii Ras Sidi-el-Ahbed (Bocoia), 100 m. alt.; 1 febr.” Lectotype (designated here):—In umbrosis maritimis promontorii Ras Sidi-el-Ahbed (Bocoia), 100 m, 1 February 1929, Font Quer 449-1929 (lectotype BC 98253! [https://www.ibb.csic.es/herbari/ JPEG / BC 98253.jpg]; isolectotypes BCN 72591 [photo!], GDA 5360 [photo!], MA 130099!, MPU 006647 [photo!]) Notes:— Carrasco (1975) cites this taxon as Senecio leucanthemifolius var. latisectus Pau & Font Quer. The GDA specimen was labelled as syntype by Vizoso in 2013. The BC specimen was identified as S. leucanthemifolius Poir. by Blanca in 1996, and the MPU voucher as S. leucanthemifolius var. fradinii (Pomel) Maire (= subsp. mauritanicus ) by Alexander in 1975.

1 Senecio foliosus var. paludicolus Pau in Cavanillesia 1: 142 (1929a)

= Jacobaea vulgaris Gaertn.

Ind. loc.:—“Pantano de Negrón (Vidal)”

Type:—not seen

Senecio nebrodensis var. occidentalis Pau , in sched. ( BC 89909! [https://www.ibb.csic.es/herbari/ JPEG / BC 89909. jpg]), nom. nud.

= Senecio squalidus L.

Note:—The specimen was labelled by Pau as “var. n.” and identified as S. squalidus by Blanca in 1996.

1,2,3 Senecio perralderianus var. hosmariensis Pau & Font Quer in Font Quer, Iter Marocc. 1928 nº 405 (1929), in sched.

= Senecio perralderianus Coss. & Durieu

Ind. loc.:—“Hab. in quercetis ilicis montis Kelti (Yebala), solo calc., 1500 m. alt.; 18 junii”

Lectotype (designated by Calvo et al. 2015: 76):—In quercetis ilicis montis Kelti (Yebala), solo calc., 1500 m, 18 June 1928, Font Quer 405-1928 (lectotype MA 130562 [photo!] [http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00130562]; isolectotypes BC 29442 !*, BCN 71366 [photo!], EMMA 18549 [photo!], G 00308028 [photo!]*, GDA 5436 [photo!], MPU 006599 [photo!]*)

Note:—The MA specimen was labelled as lectotype by Calvo in 2012. The G and GDA specimens were labelled as isolectotypes by Calvo in 2012 and Vizoso in 2016, respectively. The BC, G, and MPU specimens were cited by Calvo et al. (2015) as isolectotypes. The GDA specimen was identified as S. perralderianus subsp. hosmariensis (Pau & Font Quer) Blanca by Blanca in 1996, and the G and MA vouchers as S. perralderianus by Calvo in 2012.

1,2,3 Sideritis arborescens var. africana Font Quer & Pau in Font Quer, Iter Marocc. 1927 nº 532 (1928), in sched.

= Sideritis arborescens subsp. maireana (Font Quer & Pau) Socorro & Arreb.

Ind. loc.:—“ Hab. in saxosis calc. littoris rhiphaei, pr. Torres de Alcalá (Beni Bu-Frah), 100 m. alt.; 4 junii” Lectotype (designated by Obón & Rivera 1994: 454):—In saxosis calc. littoris rhiphaei, pr. Torres de Alcalá (Beni Bu-Frah), 100 m, 4 June 1927, Font Quer 532-1927 (lectotype UC 370029 [photo!] [https://webapps.cspace. berkeley.edu/ucjeps/object/ UC370029 ]; isolectotypes BC 74187 !, BCN 18033 [photo!], G 00424807 [photo!], GDA 32524 [photo!], MA 100746 !, MPU 006265 [photo!])

Note:— The UC specimen was labelled as lectotype by Rivera & Obón in 1992 and the GDA specimen as isolectotype by Vizoso in 2013 .

1,2,3 Sideritis arborescens var. ortonedae Font Quer & Pau in Font Quer, Cavanillesia 3: 60 (1930b)

≡ Sideritis arborescens subsp. ortonedae (Font Quer & Pau) Maire

Ind. loc.:—“Hab. cum praecedentibus, in monte Cudia Arneb, dicto”

Lectotype (designated by Obón & Rivera 1994: 550):—In collibus arenaceo-calc. Cudia Arneb (Beni-Bu-Iahi), 550 m, 29 May 1929, Font Quer 373-1929 (lectotype UC 420142 [photo!] [https://webapps.cspace.berkeley. edu/ucjeps/object/ UC420142 ]; isolectotypes BC 97956 !, BCN 72518 [photo!], G 00424810 [photo!], GDA 54787 [photo!], MA 100726 !)

Notes:—The UC specimen was labelled as lectotype by Rivera & Obón in 1992, and the GDA specimen as isolectotype by Vizoso in 2013. El Oualidi et al. (2012) mention as types two sheets at Herb. MPU ( MPU 001254 [photo!], MPU 001255 [photo!]) collected in 1932 by Sennen & Mauricio in Afso, which can not be considered original material as it was collected after publication of the name by Font Quer (1930b).

1,2,3 Sideritis briquetiana Font Quer & Pau in Font Quer, Cavanillesia 3: 60 (1930b)

Ind. loc.:—“ Hab. in declivibus calc. montis Cudia Arneb dictis (Beni-Bu-Iahi), Imp. Maroc., ad 550 m. alt., ubi, d. 29 majii 1929”

Lectotype (designated by Obón & Rivera 1994: 489):—In collibus arenaceo-calcareis Cudia Arneb dict. (Beni-Bu-Iahi), 550 m, 29 May 1929, Font Quer 375-1929 (lectotype UC 420144 [photo!] [https://webapps.cspace. berkeley.edu/ucjeps/object/ UC420144 ]; isolectotypes BC 97958 !, BCN 72208 [photo!], G 00424815 [photo!], MPU 008000 [photo!])

Notes:— APD (2020) cites this taxon as S. briquetiana Font Quer , but according to the protologue the authority should be attributed to Font Quer & Pau. Determination of the UC specimen was confirmed by Obón & Rivera in 1992.

1,3 Sideritis ×candelii Font Quer & Pau in Font Quer, Cavanillesia 3: 61 (1930b) [ Sideritis arborescens subsp. ortonedae (Font Quer & Pau) Maire × S. briquetiana Font Quer & Pau ]

Ind. loc.:—“Hab. cum praecedentibus, in monte Cudia Arneb”

Lectotype (designated here):—In collibus Cudia Arneb dictis (Beni Bu-Iahi), 550 m, 29 May 1929, Font Quer

377-1929 ( lectotype BC 97960 ! [https://www.ibb.csic.es/herbari/ JPEG / BC97960 .jpg]; isolectotypes BC 836244 !,

BCN 72207 [photo!], GDA 32540 [photo!], MA 101105!, MPU 006657 [photo!])

Notes:— The GDA specimen was labelled as sytnype by Vizoso in 2013. The BC specimen selected as lectotype is the one with more abundant samples .

1,2,3 Sideritis maireana Font Quer & Pau in Font Quer, Iter Marocc. 1927 nº 533 (1928), in sched.

≡ Sideritis arborescens subsp. maireana (Font Quer & Pau) Socorro & Arreb.

Ind. loc.:—“Hab. in saxosis arenaceis supra Bu-Meziat (Atlante rhiphaeo), 1700 m. alt.; 29 junii”

Lectotype (designated by Obón & Rivera 1994: 466):—In saxosis arenaceis supra Bu-Meziat (Atlante rhiphaeo), 1700 m, 29 June 1927, Font Quer 533-1927 (lectotype UC 370030 [photo!] [https://webapps.cspace.berkeley. edu/ucjeps/object/ UC370030 ]; isolectotypes BC 74312 !, BCN 18032 [photo!], G 00424908 [photo!], GDA 54795 [photo!], MA 100748 !, MPU 006267 [photo!], MPU 006266 [photo!])

Notes:—The GDA specimen was labelled as isolectotype by Vizoso in 2013, and the UC voucher as lectotype by Rivera & Obón in 1992. The MA specimen was identified as S. arborescens Salzm. ex Benth. by Morales in 2006. Rejdali & Fennane (2007) do not accept subspecies in S. arborescens , and include S. maireana in the synonymy of this species, although Rejdali (2002b) previously accepted S. maireana at the species rank. The main difference between these taxa is the size of the lower verticillaster, up to 12 mm wide in S. arborescens , and more than 15 mm in S. maireana .

1,2,3 Sideritis subatlantica var. rhiphaea Font Quer & Pau in Font Quer, Iter Marocc. 1927 nº 530 (1928), in sched.

= Sideritis subatlantica Doum. ex Batt.

Ind. loc.:—“Hab. in declivibus calc. montis Yebel Hammam (Littoris rhiphaei), 200 m. alt.; 16 majii” Lectotype (designated by Peris et al. 1995: 2):—In declivibus calc. montis Yebel Hammam (Littoris rhiphaei), 200 m, 16 May 1927, Font Quer 530-1927 (lectotype BC 74305! [https://www.ibb.csic.es/herbari/ JPEG / BC 74305. jpg]; isolectotypes BCN 18034 [photo!], GDA 32597 [photo!], MA 100736!, MPU 006655 [photo!]) Notes:— APD (2020) attributes this variety only to Font Quer, but on the label both authors are cited. The GDA specimen was labelled as syntype by Vizoso in 2013.

1,2,3 Silene cuatrecasasii Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 190 (1928), in sched.

Ind. loc.:—“Hab. in saxosis arenaceis ad pedem montis Yebel Sidel dictis (Atlante rhiphaeo), 1700 m. alt.; 3 julii”

Lectotype (designated here):—In saxosis arenaceis ad pedem montis Yebel Sidel dictis (Atlante rhiphaeo), 1700 m, 3 July 1927, Font Quer 190-1927 (lectotype BC 9074 ! [https://www.ibb.csic.es/herbari/ JPEG / BC9074 .jpg]; isolectotypes BC 811123 !, BCN 17295 [photo!], BM 000593501 [photo!], GDA 2167 [photo!], MA 31649 !, MA 31651 !, MPU 006343 [photo!], S-G 8712 [photo!])

Notes:— The GDA specimen was labelled as syntype by Vizoso in 2013, and the MPU voucher as isotype by Sennedot and Scouppe in 2006. The chosen lectotype is the BC specimen with the original label of the exsiccata Iter Maroccanum . Determination of BC 811123 ! was confirmed by Montserrat in 2005 .

1,2,3 Silene divaricata var. brachycalyx Pau & Font Quer in Font Quer, Iter Marocc. 1929 nº 145 (1930a), in sched.

= Silene aellenii Sennen

Ind. loc.:—“Hab. in saxosis calc. montis Targsunt (Bocoia), ad 350 m. alt.; 16 majii”

Lectotype (designated here):—In saxosis calc. montis Targsunt (Bocoia), 350 m, 16 May 1929, Font Quer 145- 1929 (lectotype BC 98196 ! [https://www.ibb.csic.es/herbari/ JPEG / BC98196 .jpg]; isolectotypes BCN 72236 [photo!], GDA 2177 , MA 31926 !, MPU 006696 [photo!])

Notes:— The GDA specimen was labelled as syntype by Vizoso in 2013. The BC specimen was identified as S. aellenii by Talavera in 1996 .

1,2,3 Silene grosiana Pau & Font Quer in Font Quer, Iter Marocc. 1928 nº 91 (1929), in sched.

= Silene ibosii Emb. & Maire

Ind. loc.:—“ Hab. in arenosis, c. Xauen, 600 m. alt.; 25 aprilis”

Lectotype (designated by Fernández Casas 1982: 9):—in arenosis, c. Xauen, 600 m, 25 April 1928, Font Quer 91- 1928 (lectotype MA 31809 ! [http://colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00031809]; isolectotypes BC 9079 !, BCN 17955 [photo!], BM 000990894 [photo!], GDA 2209 [photo!], MPU 006435 [photo!], MPU 006436 [photo!], Z 000002681 [photo!])

Notes:— The GDA specimen was labelled as syntype by Vizoso in 2013. The BC and BM specimens were identified as S. ibosii by Mateos in 1999, and by Oxelman in 2005, respectively .

1,2,3 Silene guedirensis Pau in Font Quer, Iter Marocc. 1930 nº 193 (1932), in sched.

= Silene micropetala Lag.

Ind. loc.:—“Hab. inter chamaeropes, l. El Guedira dicto, pr. El Araix; 26 aprilis”

Lectotype (designated by Talavera & Muñoz Garmendia 1989: 419):— El Guedira dicto, pr. El Araix, 26April 1930, Font Quer 193-1930 (lectotype MA 31017 ! [http://colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00031017]; isolectotypes BC 136880 !, BCN 72955 [photo!], BM 000552459 [photo!], GDA 2223 [photo!]*, MPU 006777 [photo!], S-G 8706 [photo!], Z 000002682 [photo!])

Notes:— The MA specimen was labelled as lectotype by Talavera in 1985, and the GDA voucher as isolectotype by Vizoso in 2015. Talavera & Muñoz Garmendia (1989) cite as isolectotype the GDA specimen. The BC and MA specimens were identified as S. micropetala by Talavera in 1985 .

1,2 Silene maurisca Pau in Bol. Soc. Iber. Ci. Nat. 31: 96 (1934)

= Silene patula Desf. subsp. patula

Ind. loc.:—“ Zucar [Zaccar] de Miliana, localidad de Battandier y Trabut (Alleizette). Atlas Medio (Jahandiez)” Lectotype (designated here):— Moyen Atlas: Aïn Leuh, bois des coteaux calcaries, 1600 m, 20 May 1924, Jahandiez 370 (lectotype MA 32118 [photo!] [http://colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00032118]; isolectotype P 04966493 [photo!])

Notes:— Two localities are indicated in the protologue: one from Algeria prospected by Alleizette ( MA 32161 !), and the other from Morocco, by Jahandiez ( MA 32118 and P 04966493 ). Specimens from Morocco are better preserved than the one from Algeria. Among the Moroccan material, the selected lectotype is the one from Herb. MA. The MA and P specimens were identified as S. patula subsp. patula by Jeanmonod in 1983 and 1985, respectively. The accepted name follows Talavera (2002b) .

1,2,3 Silene rhiphaena Pau & Font Quer in Font Quer, Iter Marocc. 1929 nº 144 (1930a), in sched.

Ind. loc.:—“Hab. in saxosis calc. supra El Zaio (Ulad Setut), ad 400 m. alt.; 11 aprilis”

Lectotype (designated here):— In saxosis calc. supra El Zaio (Ulad Setut), 400 m, 11 April 1929, Font Quer 144-1929 (lectotype BC 98195 ! [https://www.ibb.csic.es/herbari/ JPEG / BC98195 .jpg]; isolectotypes BCN 20114 [photo!], BM 000843948 [photo!], GDA 2346 [photo!], MA 31100 !, MA 31718 !, MPU 006697 [photo!], MPU 006698 [photo!], RAB 014987 [photo!]*, S-G 8694 [photo!])

Notes:— The GDA specimen was labelled as syntype by Vizoso in 2013. El Oualidi et al. (2012) mentioned the specimen at Herb. RAB as type .

1,3 Silene setacea var. microsperma Pau in Cavanillesia 4: 157 (1931)

= Silene vivianii Steud. subsp. vivianii

Ind. loc.:—“-”

Lectotype (designated here):—Sous: Sud de Tiznit , bords sablonneux de l’Assif Ouaraben, 31 March 1931, Jahandiez 82 (lectotype MA 31179 ! [http://colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01- 00031179]; isolectotype MARS 15930 [photo!])

Note:— Pau (1931) described this variety on a specimen distributed by Jahandiez in the exsiccata Plantes Marocaines 1931 nº 82, which was labelled Silene setacea .

1,2,3 Silene venosa var. rhiphaea Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 178 (1928), in sched.

= Silene vulgaris subsp. commutata (Guss.) Hayek

Ind. loc.:—“Hab. in saxosis, c. Badú (Atlante rhiphaeo), 1600 m. alt.; 4 julii”

Lectotype (designated here):—In saxosis, c. Badú (Atlante rhiphaeo), 1600 m, 4 July 1927, Font Quer 178-1927 (lectotype BC 8377 ! [https://www.ibb.csic.es/herbari/ JPEG / BC8377 .jpg]; isolectotypes BCN 16268 [photo!], GDA 2337 [photo!], MA 30398 !, MPU 006344 [photo!])

Notes:— The GDA specimen was labelled as syntype by Vizoso in 2013, and the MPU voucher as isotype by Sennedot and Scouppe in 2006. The BC specimen was identified as S. vulgaris subsp. commutata by Talavera in 1996 .

1,2,3 Silene vidaliana Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 181 (1928), in sched.

Ind. loc.:—“Hab. in arenosis, supra Bu-Meziat (Atlante rhiphaeo), 1750 m. alt.; 16 junii”

Lectotype (designated here):—In arenosis, supra Bu-Meziat (Atlante rhiphaeo), 1750 m, 16 June 1927, Font Quer

181-1927 (lectotype BC 9089! [https://www.ibb.csic.es/herbari/JPEG/BC9089.jpg]; isolectotypes BCN 17296 [photo!], G 00015455 [photo!], GDA 2325 [photo!], MA 31407!, MPU 006345 [photo!], S-G 8686 [photo!]) Note:—The GDA specimen was labelled as syntype by Vizoso in 2013, and the MPU voucher as isotype by Sennedot and Scouppe in 2006 .

1,2,3 Sonchus masguindalii Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 732 (1928), in sched.

Ind. loc.:—“ Hab. in fissuris rupium calc. littoris rhiphaei, c. Marsa Quebira (Bocoya), 20 m. alt.; 18 majii” Lectotype (designated here):—In fissuris rupium calc. littoris rhiphaei, c. Marsa Quebira (Bocoya), 20 m, 18 May 1927, Font Quer 732-1927 (lectotype MA 139928 ! [http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00139928]; isolectotypes BC 35712 !, BCN 17963 [photo!], G 00018358 [photo!], GDA 5543 [photo!], MPU 006363 [photo!], MPU 006364 [photo!])

Notes:— The BC specimen includes poor and not well conserved material, therefore the MA specimen is selected as lectotype. Determination of the BC specimen was confirmed by Mejías in 1995 .