Narcissus bulbocodium f. auratus Pau

Nualart, Neus, Soriano, Ignasi, Prieto, David Pérez & Ibáñez, Neus, 2021, Catalogue and typification of the Moroccan taxa published by Carlos Pau, Phytotaxa 519 (1), pp. 1-94 : 56-62

Narcissus bulbocodium f. auratus Pau in Font Quer, Iter Marocc. 1929 nº 93 (1930a), in sched., nom. nud.

= Narcissus romieuxii Braun-Blanq. & Maire

Notes:—The form name was published under “ N. bulbocodium var. candicans (Haw.) Pau ” and distributed in the exsiccata Iter Maroccanum 1929 without a description. Maire (1931) identified this form as N. bulbocodium subsp. romieuxii var. rifanus Emb. & Maire. A specimen of this plant (MA 148137!) was identified as N. cantabricus DC. by Aedo in 2008, but another (GDA 215 [photo!]) as N. romieuxii subsp. romieuxii var. rifanus (Emb. & Maire) A.Fern. by Fernández-Casas in 2014. The name “auratus” proposed by Pau seems more appropriate for N. romieuxii (with sulphur flowers) than for N. cantabricus (with white flowers). Anyway, all these infraspecific taxa are included by APD (2020) in N. romieuxii .

1,2,3 Nonea perezii Pau in Bol. Soc. Esp. Hist. Nat. 14: 428 (1914)

= Nonea heterostemon Murb.

Ind. loc.:—“alrededores de Larache … Sr. Pérez Camarero, durante la primavera pasada [1914]”

Lectotype (designated by Nualart et al. 2018: 614):—Larache (Afr. bor), 1914, Pérez Camarero 34 (lectotype BC 42193 ! [https://www.ibb.csic.es/herbari/ JPEG / BC42193 .jpg]; isolectotype BC-Sennen 990170!)

1,2,3 Oenanthe maroccana Pau & Font Quer in Font Quer, Iter Marocc. 1928 nº 297 (1929), in sched.

=§ Oenanthe crocata L.

Ind. loc.:—“Hab. in paludosis, ad pedem Yebel Afestal (Gomara), 1500 m. alt.”

Lectotype (designated here):—In paludosis, ad pedem Yebel Afestal (Gomara), 1500 m, 25 June 1928, Font Quer 297-1928 (lectotype BC 25087 ! [https://www.ibb.csic.es/herbari/ JPEG / BC25087 .jpg]; isolectotypes BCN 17837 [photo!], GDA 40605 [photo!], MA 87963 !, MA 87964 !, MPU 006622 [photo!], MPU 006623 [photo!], Z 000028709 [photo!])

Notes:—The GDA specimen was labelled as syntype by Vizoso in 2013.The BC and MA specimens were identified as O. crocata by Jury in 1997 and Southam in 2002, respectively. APD (2020) synonymizes this species under O. pimpinelloides subsp. callosa (Salzm. ex DC.) Maire , but Southam (2002) synonymized it under O. crocata . The original material of O. maroccana shows rays and pedicels not thickened in fruit, a feature that matches O. crocata according to Southam (2002) and El Alaoui Faris & Ibn Tattou (2007).

1,3 Onobrychis cadevallii Jahand., Maire & Pau in Jahandiez & Maire, Bull. Soc. Hist. Nat. Afrique N. 19(2): 83 (1928) Ind. loc.:—“Hab. in pascuis rupestribus calcareis, inter cedreta Atlantis Medii , ad alt. 1700–2300 m, junio florens: in monte Koubbat prope Bekrit (JAHANDIEZ, 1924, n° 549); in monte Hayan; ad fontes amnis Ifrane” Lectotype (designated here):— Moyen Atlas : Bekrit , foret de Cedres de Koubbat, 1950 m, 22 June 1924, Jahandiez 549 (lectotype MPU 000337 [photo!]* [https://herbier.umontpellier.fr/hv/wakka.php?wiki=PagePrincipale/ifram e&action=detail&ordre=p&codebarre= MPU000337 ]; isolectotypes P 00084445 [photo!])

Notes:— This species was published by Jahandiez & Maire (1928). El Oulaidi et al. (2012) mention three syntypes for this species: MPU 000337 from Bekrit, MPU 000338 [photo!] from Ifrane, and MPU 000339 [photo!] from Hayan. It should be noted that there are two gatherings with number 549 from Bekrit considered both as syntypes: plants collected on 5 June ( BC 143024 !, MA 343995 !) and plants collected on 22 June ( MPU 000337 , P 00084445 ). The specimen of the Maire collection in Herb. MPU, labelled as type by Maire himself, is selected as lectotype. The MPU specimen from Ifrane and the P specimen were labelled as syntypes by Durand and Loup and Cottin respectively, both in 2004 .

1,2,3 Onobrychis ligulifera Pau in Ann. Sci. Acad. Polytechn. Porto 6: 99 (1911)

= Onobrychis crista-galli (L.) Lam.

Ind. loc.:—“ Melilla … En sitios incultos”

Holotype:— Corazón de Toro en el barranco que se cruza para venir a Zeluan, 9 May 1910, Pau s.n. ( MA 68933 ! [http://colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00068933])

3 Onobrychis peduncularis var. maura Pau ex Jahandiez & Maire in Bull. Soc. Hist. Nat. Afrique N. 19(2): 84 (1928) = Onobrychis humilis (L.) G.López subsp. humilis

Ind. loc.:—“ Hab. in pascuis, in calvitiis quercetorum Atlantis Medii , solo calcareo: Aïn-Leuh, ad alt. 1400–1500 m (JAHANDIEZ, 1924, n° 285); Daya Chiker, ad alt. 1400 m (HUMBERT et MAIRE, 1925). - Maio floret - Typus in Herb. Univers. Algeriensis, in Herb. Inst. Imper. Scient. Rabatensis, in Herb. JAHANDIEZ et in Herb. C. PAU”

Lectotype (designated here):— In Atlantis Medii montibus supra urbem Taza: Daya Chiker in calviticis quercetorum, 19 June 1925, Maire s.n. ( MPU 000341 [photo!] [https://herbier.umontpellier.fr/hv/wakka. php?wiki=PagePrincipale/iframe&action=detail&ordre=p&codebarre= MPU000341 ])

Notes:—There are several syntypes of this taxon: MA 68993! and MPU 000340 [photo!] from Aïn-Leuh, and MPU 000341 [photo!]), from Daya Chiker. The selected lectotype is the only one with mature fruits in the Herb. MPU. It should be noted that the MA specimen has a revision label by Valdés in 1999 indicating that both samples in this sheet belong to two different taxa: O. humilis subsp. humilis and O. africana Sirj. The taxonomy of this last species is not clear: APD (2020) includes it in O. conferta (Desf.) Desv. subsp. conferta , but El Oualidi & Navarro (2007a) in O. cadevallii Jahand., Maire & Pau. The plants of the MA specimen, once examined, both correspond to Onobrychis humilis subsp. humilis .

1,2,3 Ononis arborescens var. glabrescens Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 290 (1928), in sched.

= Ononis hispida subsp. arborescens (Desf.) Sirj.

Ind. loc.:—“Hab. in saxosis arenaceis, pr. Badú (Atlante rhiphaeo), 1600 m. alt.; 6 julii”

Lectotype (designated here):—In saxosis arenaceis, pr. Badú (Atlante rhiphaeo), 1600 m, 6 July 1928, Font Quer 290-1928 (lectotype BC 14788 ! [https://www.ibb.csic.es/herbari/ JPEG / BC14788 .jpg]; isolectotypes BCN 17307 [photo!], GDA 34356 [photo!], MA 60997 !, MPU 006306 [photo!])

Notes:— Carrasco (1975) and Devesa (2002a) cite this taxon as Ononis hispida var. glabrescens Pau & Font Quer. The MPU and GDA specimens were labelled as syntypes by Mathez in 1982 and Vizoso in 2013, respectively.

1,2,3 Ononis arborescens var. remotiflora Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 289 (1928), in sched.

= Ononis hispida subsp. arborescens (Desf.) Sirj.

Ind. loc.:—“Hab. in cistetis, supra Targuist, 1100 m. alt., solo siliceo, fl. 5; fruct. 29 julii”

Lectotype (designated here):—In cistetis, supra Targuist, 1100 m, fl. 5 June 1927, fruct. 29 June 1927, Font Quer 289-1927 (lectotype BC 14787 ! [https://www.ibb.csic.es/herbari/ JPEG / BC14787 .jpg]; isolectotypes BM 000843201 [photo!], BCN 17308 [photo!], GDA 34355 [photo!], MA 60999 , MPU 006308 [photo!], MPU 006309 [photo!])

Notes:— Carrasco (1975) cites this taxon as Ononis hispida subsp. arborescens var. remotiflora Pau & Font Quer. The MPU and GDA specimens were labelled as syntypes by Mathez in 1982 and Vizoso in 2013, respectively.

1,2,3 Ononis cephalotes var. minutifolia Pau & Font Quer in Font Quer, Cavanillesia 4: 65 (1931d)

= Ononis cephalotes Boiss.

Ind. loc.:—“Hab. in monte Dj. Tazaut (Beni Zedjel), ad 1750 m. alt., ubi, d. 9 julii 1930”

Lectotype (designated here):— In collibus calc. montis Dj. Tazaut (Beni Zedjel), 1750 m, 9 July 1930, Font Quer 329-1930 (lectotype BC 98134 ! [https://www.ibb.csic.es/herbari/ JPEG / BC98134 .jpg]; isolectotypes GDA 34379 [photo!], MA 61112 !, MA 61113 !, MPU 007144 [photo!])

Note:—The original label of the exsiccata Iter Maroccanum 1930 includes the mention “cotypus” (= syntype).

1,2,3 Ononis cintrana var. macrodonta Pau in Font Quer, Iter Marocc. 1930 nº 321 (1932), in sched.

= Ononis cintrana Brot.

Ind. loc.:—“Hab. in monte Dj. Quebir, c. Tandja; 16 aprilis”

Lectotype (designated here):— In monte Dj. Quebir, c. Tandja, 16 April 1930, Font Quer 321-1930 (lectotype BC 98126 ! [https://www.ibb.csic.es/herbari/ JPEG / BC98126 .jpg]; isolectotypes BCN 73166 [photo!], GDA 34384 [photo!], MA 61310 , MPU 006745 [photo!])

Note:— The GDA specimen was labelled as syntype by Vizoso in 2013 .

2,3 Ononis inaequifolia var. minoriflora Pau in Font Quer, Iter Marocc. 1927 nº 296 (1928), in sched.

= Ononis natrix subsp. arganietorum (Maire) Sirj.

Ind. loc.:—“Hab. in incultis, c. Targuist, 1000 m. alt.; 1 junii”

Lectotype (designated here):—In incultis, c. Targuist, 1000 m, 1 June 1927, Font Quer 296-1927 (lectotype BC 14523 ! [https://www.ibb.csic.es/herbari/ JPEG / BC14523 .jpg]; isolectotypes BCN 17306 [photo!], GDA 34404 [photo!], MA 61385 !, MPU 006305 [photo!])

Notes:—The GDA specimen was labelled as syntype by Vizoso in 2013. The BC specimen was identified as Ononis natrix subsp. arganietorum by Devesa in 1994.

1,2,3 Ononis maweana var. fontqueri Pau in Bol. Soc. Esp. Hist. Nat. 14: 427 (1914)

= Ononis maweana Ball

Ind. loc.:—“alrededores de Larache … Sr. Pérez Camarero, durante la primavera pasada [1914]”

Lectotype (designated by Nualart et al. 2018: 616):—Larache, 1914, Pérez Camarero 15 (lectotype MA 61795! [http://colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00061795]; isolectotype BC 14553!)

1,3 Ononis pendula var. grandiflora Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 311 (1924a), nom. illeg.

=§ Ononis pendula subsp. boissieri (Sirj.) Devesa

Ind. loc.:—“En las cercanías de Tetuán y en las faldas del Dersa”

Holotype:—Tetuan, May 1921, Pau s.n. ( MA 61595 [photo!] [http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00061595])

Notes:—This name should be considered illegitimate according to Art. 53.1 ( Turland et al. 2018) since a homonym exists that was published by Boissier (1840: 153). APD (2020) includes this variety in O. pendula subsp. munbyi (Sirj.) Greuter & Burdet , but according to the description by Pau (1924a) (corolla of 20 mm and calyx teeth wide) and the revision of the holotype by Devesa in 1996, it should be synonymized under O. pendula subsp. boissieri . There are other specimens labelled as O. pendula var. grandiflora Pau and distributed in Font Quer’s exsiccata Iter Maroccanum 1928 nº 185 that do not match the description by Pau (1924a) (the corolla size does not exceed 13 mm) and therefore should be identified as O. pendula Desf. subsp. pendula .

§ 1,2,3 Ononis penduliflora Pau in Font Quer, Iter Marocc. 1930 nº 325 (1932), in sched.

Ind. loc.:—“Hab. in arenosis, pr. El Araix; 16 aprilis”

Lectotype (designated by Devesa 1996: 194):— In arenosis, pr. El Araix, 16 April 1930, Font Quer 325-1930 (lectotype MA 61053 ! [http://colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00061053]; isolectotypes BC 98130 !, BC 810359 !, BC 908900 !, BCN 73120 [photo!], BM 000843199 [photo!], GDA 34444 [photo!], MPU 006744 [photo!], Z 000023612 [photo!])

Notes:— APD (2020) synonymizes this taxon under O. pendula subsp. munbyi , but Devesa (2002a), Ibn Tattou (2007d), and Rankou et al. (2013) accept this species as a Moroccan endemic plant. Ononis pendula Desf. and O. penduliflora differ by the size of the calyx teeth and the shape of the leaflets.

1,2,3 Ononis pubescens var. semiunifolia Pau & Font Quer in Font Quer, Iter Marocc. 1928 nº 186 (1929), in sched.

= Ononis pubescens L.

Ind. loc.:—“Hab. in quercetis suberis, ad pedem Yebel Sugna (Yebala), 500 m. alt.; 26 majii”

Lectotype (designated here):—In quercetis suberis, ad pedem Yebel Sugna (Yebala), 500 m, 26 May 1928, Font Quer 186-1928 (lectotype BC 14610 ! [https://www.ibb.csic.es/herbari/ JPEG / BC14610 .jpg]; isolectotypes BCN 17672 [photo!], GDA 34455 [photo!], MA 61632 !, MPU 006407 [photo!])

Note:— The GDA specimen was labelled as syntype by Vizoso in 2013 .

1,2,3 Ononis simulata Pau & Font Quer in Font Quer, Iter Marocc. 1928 nº 182 (1929), in sched.

≡ Ononis alopecuroides subsp. simulata (Pau & Font Quer) Maire

Ind. loc.:—“Hab. in valle fl. Lau, c. Bab Taza, 600 m. alt.; solo arenoso; 2 junii”

Lectotype (designated here):—In valle fl. Lau, c. Bab Taza, 600 m, 2 June 1928, Font Quer 182-1928 (lectotype BC 14819 ! [https://www.ibb.csic.es/herbari/ JPEG / BC14819 .jpg]; isolectotypes BCN 18017 [photo!], BM 000843174 [photo!], GDA 34494 [photo!], MA 61103 !, MPU 006410 [photo!], Z 000023613 [photo!]) Notes:—The GDA specimen was labelled as syntype by Vizoso in 2013. The BC specimen was identified as O. alopecuroides subsp. simulata by Devesa in 1996.

1,2,3 Ononis tetuanensis Pau in Vidal, Bol. Soc. Esp. Hist. Nat. 21: 276 (1921)

= Ononis pendula Desf. subsp. pendula

Ind. loc.:—“Xauen, 15- V ”

Holotype:—Xauen, 15 May 1921, Vidal s.n. ( MA 61602 ! [http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00061602])

1,2,3 Ononis thomsonii var. grandiflora Pau & Font Quer in Font Quer, Iter Marocc. 1930 nº 314 (1932), in sched.

= Ononis thomsonii Oliv.

Ind. loc.:—“Hab. in saxosis calc. cacuminis montis Lexhab (Gomara), ad 2050 m. alt.; 21 julii”

Lectotype (designated here):—In saxosis calc. cacuminis montis Lexhab (Gomara), 2050 m, 21 July 1930, Font Quer 314-1930 (lectotype BC 98119 ! [https://www.ibb.csic.es/herbari/ JPEG / BC98119 .jpg]; isolectotypes BCN 73160 [photo!], GDA 34508 [photo!], MA 61819 !, MPU 006746 [photo!], VIT 32366 [photo!])

Note:— The GDA specimen was labelled as syntype by Vizoso in 2013 .

1,2,3 Ononis thomsonii var. parvifolia Pau & Font Quer in Font Quer, Iter Marocc. 1928 nº 187 (1929), in sched.

= Ononis thomsonii Oliv.

Ind. loc.:—“Hab. in glareosis calc., cacuminis Yebel Kelti, 1900 m. alt.”

Lectotype (designated here):—In glareosis calc., cacuminis Yebel Kelti, 1900 m, 18 June 1928, Font Quer 187- 1928 (lectotype BC 14762 ! [https://www.ibb.csic.es/herbari/ JPEG / BC14762 .jpg]; isolectotypes BCN 17964 [photo!], GDA 34507 [photo!], MA 61820 !, MPU 006408 [photo!])

Notes:— The MPU specimen was labelled as isotype by Sennedot & Grossmann in 2006, and the GDA specimen as syntype by Vizoso in 2013. The MA specimen was identified as O. thomsonii by Cannon in 1979 .

1,2,3 Onosma maroccana Pau in Font Quer, Iter Marocc. 1930 nº 538 (1932), in sched.

= Onosma tricerosperma subsp. mauritanica (Maire) G.López

Ind. loc.:—“Hab. in saxosis calc. montis Lexhab (Gomara), ad 2000 m. alt.; 22 julii”

Lectotype (designated here):—In saxosis calc. montis Lexhab (Gomara), 2000 m, 22 July 1930, Font Quer 538-1930 (lectotype BC 98036 ! [https://www.ibb.csic.es/herbari/ JPEG / BC98036 .jpg]; isolectotypes BCN 73382 [photo!], GDA 976 [photo!], MA 96749 !, MPU 005275 [photo!], S-G 4368 [photo!], Z 000053949 [photo!]) Notes:—The GDA specimen was labelled as syntype by Vizoso in 2013. The BC specimen was identified as O. fastigiata subsp. mauritanica Maire by Mateos in 1997.

1 Ophrys × composita Pau in Monde Pl. 30(66–181): 1 (1929b), nom. nud.

= Ophrys × peltieri Maire [ Ophrys scolopax Cav. × O. tenthredinifera Willd. ]

Note:—This name was published only with the indication of the parent species: Ophrys rosea (Desf.) Samp. × O. scolopax .

1,2,3 Orchis incarnata var. xauensis Pau & Font Quer in Font Quer, Iter Marocc. 1928 nº 56 (1929), in sched.

= Dactylorhiza durandii (Boiss. & Reut.) M.Laínz

Ind. loc.:—“Hab. in paludosis montis Kalaa, 1300 m. alt.; 15 junii”

Lectotype (designated here):—In paludosis montis Kalaa, 1300 m, 15 June 1928, Font Quer 56-1928 (lectotype BC 60462 ! [https://www.ibb.csic.es/herbari/ JPEG / BC60462 .jpg]; isolectotypes BCN 17490 [photo!], BM 000013513 [photo!], GDA 36717 [photo!], MA 24009 !, MPU 006439 [photo!], MPU 006440 [photo!], S 06- 20724 [photo!])

Notes:—The GDA specimen was labelled as syntype by Vizoso in 2013. The BC specimen was identified as Dactylorhiza elata subsp. durandii (Boiss. & Reut.) Soó by Silvestre in 1995.

§ 1,2,3 Origanum ×fontqueri Pau in Font Quer, Iter Marocc. 1930 nº 578 (1932), in sched. (“ font-queri ”) [ Origanum compactum Benth. × O. grosii Pau & Font Quer ]

Ind. loc.:—“Hab. in declivibus calc. c. Talambot, ad 650 m. alt., inter parentes”

Lectotype (designated by Ietswaart 1980: 101):—In declivibus calc. c. Talambot, 650 m, 8 July 1930, Font

Quer 578-1930 (lectotype MA 105456! [http://colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-

00105456]; isolectotypes BC 98310!, BCN 73759 [photo!], G *, MPU 005270 [photo!] (pro parte ))

Notes:— Ietswaart (1980) indicates this name as a nomen nudum although a description is provided on the label, and according to Art. 30.8 ( Turland et al. 2018) the printed labels of specimens constitute effective publications before 1 January 1953. Ietswaart (1980) also considers that the features of O. ×fontqueri do not differ enough from those of O. grosii , and synonymizes these two taxa. Rejdali (2002a) mentions O. ×fontqueri as a synonym of O. grosii , but El Oualidi & Navarro (2007b) accept O. × fontqueri as a hybrid. On the other hand, APD (2020) includes both this hybrid and O. grosii under O. elongatum (Bonnet) Emb. & Maire. According to genome size this supposed hybrid should be a result of gene flow between its parent taxa, O. grosii and O. compactum ( Bakha et al. 2017) . The G specimen was cited as isotype by Ietswaart (1980). The MA specimen was identified as Origanum grosii by Ietswaart in 1979. It should be noted that the MPU sheet preserves two different plants, the one on the left should be O. grosii , and the one on the right, O. × fontqueri .

§ 1,2,3 Origanum grosii Pau & Font Quer in Font Quer, Iter Marocc. 1928 nº 352 (1929), in sched.

Ind. loc.:—“Hab. in saxosis calc. montis Kalaa, 1000 m. alt.; 29 junii”

Lectotype (designated by Ietswaart 1980: 101):—In saxosis calc. montis Kalaa, 1000 m, 29 June 1928, Font Quer 352-1928 (lectotype MA 105455 ! [http://colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01- 00105455]; isolectotypes BC 50129 ! (pro parte), BCN 71180 [photo!], G 00424488 [photo!]*, MPU *, S-G 4383 [photo!], Z 000018959 [photo!])

Notes:—Determination of the G, MA, and S specimens were confirmed by Ietswaart in 1979. The G and MPU specimens were mentioned as isotypes by Ietswaart (1980) and Dobignard (2009), respectively. The BC sheet includes two different plants, the three samples on the left are Origanum grosii , but the one on the right is O. × fontqueri . Ietswaart (1980) considers O. grosii invalidly published as in the case of O. × fontqueri , but according to Art. 30.8 ( Turland et al. 2018) the description included in the label is sufficient to validate the name. APD (2020) synonymizes O. grosii under O. elongatum (Bonnet) Emb. & Maire , but according to Rejdali (2002a) and El Oualidi & Navarro (2007b) they can be considered different taxa, due to the size of the spikes and the pubescence of the leaves.

1 Ornithopus compositus Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 316 (1924a), nom. nud., pro hyb.

= Ornithopus compressus L.

Notes:—In the protologue there is only an indication of the parent species, Ornithopus compressus L. × O. isthmocarpus Coss. The accepted name follows Talavera & Arista (2000).

1,2,3 Paeonia mascula var. maroccana Pau & Font Quer in Font Quer, Iter Marocc. 1928 nº 105 (1929), in sched.

= Paeonia mascula subsp. coriacea (Boiss.) Malag.

Ind. loc.:—“Hab. in silvis pinsaporum Yebel Tisuka, 1800 m. alt.; 13 junii”

Lectotype (designated by Romo 1992: 207):—In silvis pinsaporum Yebel Tisuka, 1800 m, 13 June 1928, Font Quer 105-1928 (lectotype BC 1921 ! [https://www.ibb.csic.es/herbari/ JPEG / BC1921 .jpg]; isolectotypes BCN 18024 [photo!], G 00383192 [photo!], GDA 36838 [photo!, MA 38481 !, MA 38482 !, MPU 006432 [photo!], MPU 006433 [photo!])

Notes:— Carrasco (1975) cites this taxon as “ Paeonia corallina subsp. coriacea var. maroccana Pau & Font Quer ”. The G and BC specimens were identified as P. coriacea Boiss. by Hong in 1991 and Valdés in 1995, respectively.

1,2,3 Papaver rupifragum var. maroccanum Font Quer & Pau in Font Quer, Iter Marocc. 1930 nº 223 (1932), in sched. = Papaver rupifragum Boiss. & Reut.

Ind. loc.:—“Hab. in saxosis calc. montis Lexhab (Gomara), ad 2050 m. alt.; 22 julii”

Lectotype (designated here):—In saxosis calc. montis Lexhab (Gomara), 2050 m, 22 July 1930, Font Quer 223- 1930 (lectotype BC 137382 ! [https://www.ibb.csic.es/herbari/ JPEG / BC137382 .jpg]; isolectotypes B 10 0160639 [photo!]*, BCN 72973 [photo!], BM *, G 00014173 [photo!], GDA 36566 [photo!], MA 42740 !, MA 42741 !, MPU 006774 [photo!], S-G 4541 [photo!])

Notes:—Several of the above mentioned specimens were identified as P. rupifragum : the B specimen by Kadereit in 1992, the BC specimen by Valdés in 1995, and the MA specimen by Díaz González in 1983. Kadereit (1996) mentions as types the B and BM specimens, and El Oualidi et al. (2012) just cites the B one.

1 Passerina hirsuta var. oscilans Pau & Vidal in Vidal, Asoc. Esp. Progr. Ci. Congr. Cádiz 6: 158 (1928)

= Thymelaea lythroides Barratte & Murb.

Ind. loc.:—“Negrón”

Holotype:—Negrón (Hifhac), 3 m, 4 April 1926, Vidal 214 ( MA 83212 ! [http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00083212])

Note:—The specimen was labelled as lectotype by Pedrol in 1994, who identified it as Thymelaea lythroides .

3 Peplis australis [unranked] rhiphatlantica Font Quer & Pau in Font Quer, Iter Marocc. 1927 nº 424 (1928), in sched., nom. nud.

= Lythrum portula ( L.) D. A.Webb.

Notes:—The name was published as “fma. vel var.” in a label without diagnosis nor description. APD (2020) cites this name at the varietal rank.

2,3 Phalangium algeriense [unranked] brachylepis Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 115 (1928), in sched.

= Anthericum baeticum (Boiss.) Boiss.

Ind. loc.:—“ Hab. in declivibus calc. c. Torres de Alcalá (Littore rhiphaeo), solo calc., 100 m. alt.; 4 junii” Lectotype (designated here):—in declivibus calc. c. Torres de Alcalá (Littore rhiphaeo), 100 m, 4 June 1927, Font Quer 115-1927 (lectotype S 06-3148 [photo!] [http://herbarium.nrm.se/specimens/ S06-3148 ]; isolectotypes BCN 120026 [photo!], BCN 15746 [photo!], MPU 006353 [photo!], UC 369712 [photo!])

Notes:— The name was originally published as “ var. vel fma”. The MPU specimen was labelled as isotype by Sennedot & Scouppe in 2006. The lectotype is the specimen with more flowers and includes a piece of rhizome .

1,2,3 Phalangium baeticum var. rhiphaeum Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 116 (1928), in sched. = Anthericum baeticum (Boiss.) Boiss.

Ind. loc.:—“Hab. in pratis humidis montis Yebel Lerz (Atlante rhiphaeo), 2200 m. alt.; 12 junii”

Lectotype (designated here):—In pratis humidis montis Yebel Lerz (Atlante rhiphaeo), 2200 m, 12 June 1927, Font Quer 116-1927 (lectotype MA 20361 ! [http://colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00020361]; isolectotypes BCN 15747 [photo!], GDA 37623 [photo!], MPU 006352 [photo!], UC 369713 [photo!])

Notes:— Carrasco (1975) records this taxon as Phalangium liliago var. rifeum Pau & Font Quer. The GDA was labelled as syntype by Vizoso in 2013, and the MPU as isotype by Sennedot and Scouppe in 2006. The selected lectotype is the specimen with more samples.

1,2,3 Phlomis caballeroi Pau in Bol. Soc. Aragonesa Ci. Nat. 17: 132 (1918)

≡ Phlomis purpurea subsp. caballeroi (Pau) Rivas Mart.

Ind. loc.:—“ Casablanca; 6 Junio 1915 A. Caballero”

Holotype:—Casablanca Riff., 6 June, Caballero s.n. ( MA 242065 !)

Note:—The specimen was only labelled as Phlomis L. by Pau, and identified as P. caballeroi by Velayos in 1990.

1,3 Phlomis caballeroi var. submontana Pau & Font Quer in Font Quer, Iter Marocc. 1929 nº 384 (1930a), in sched.

= Phlomis purpurea subsp. caballeroi (Pau) Rivas Mart.

Ind. loc.:—“Hab. in quercetis cocciferae montis Djebel Afgal (Beni Smih), 700 m. alt.: 10 majii”

Lectotype (designated here):—In quercetis cocciferae montis Djebel Afgal (Beni Smih), 700 m, 10 May 1929, Font Quer 384-1929 (lectotype BC 97965 ! [https://www.ibb.csic.es/herbari/ JPEG / BC97965 .jpg]; isolectotypes BCN 72522 [photo!], G 00435160 [photo!], GDA 32268 [photo!], MA 102109 !, MPU 006654 [photo!]) Notes:—The GDA specimen was labelled as syntype by Vizoso in 2013. Determination of the BC specimen was confirmed by Sáez in 1997.

3 Pimpinella dichotoma var. vegeta Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 456 (1928), in sched.

= Stoibrax pomelianum (Maire) B.L.Burtt

Ind. loc.:—“Hab. in saxosis calc. littoris rhiphaei, c. Marsa Saguira (fl.) et pr. Torres Alcalá (fruct.), 50 m. alt.; fl. 13 majii, fruct. 4 junii”

Lectotype (designated here):—In saxosis calc. littoris rhiphaei, c. Marsa Saguira (fl.) et pr. Torres Alcalá (fruct.), 50 m, fl. 13 May 1927, fruct. 4 June 1927, Font Quer 456-1927 (lectotype BC 25598 ! [https://www.ibb.csic. es/herbari/ JPEG / BC25598 .jpg]; isolectotypes BCN 18037 [photo!], GDA 33089 [photo!], MA 87025 !, MPU 006288 [photo!], MPU 006289 [photo!])

Notes:—The GDA specimen was labelled as syntype by Vizoso in 2013. The BC and MA specimens were identified as Stoibrax pomelianum by Pyke in 2013 and by Jiménez Mejías in 2012, respectively.

1,3 Plantago macrorhiza var. tangerina Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 380 (1924a)

= Plantago macrorhiza Poir.

Ind. loc.:—“ Tánger ”

Holotype:— Tanger al Fondak , 2 May 1921, Pau s.n. ( MA 116017 ! [http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00116017])

1,2,3 Polygala fontqueri Pau ex Font Quer in Font Quer, Cavanillesia 4: 91 (1931c) (“ font-queri ”)

≡ Polygala rupestris subsp. fontqueri (Pau ex Font Quer) Font Quer

Ind. loc.:—“Hab. in fissuris rupium calc. l. Uad-el-Kannar (Beni Selman-Imp. Maroc), ad 200 m. alt.; 7 junii” Lectotype (designated here):—In rupibus calc. supra Uad-el-Kannar (Beni Selman), 200 m, 7 June 1930, Font Quer 408-1930 (lectotype BC 98279!* [https://www.ibb.csic.es/herbari/ JPEG / BC 98279_b.jpg]; isolectotypes BM 000843966 [photo!], GDA 35947 [photo!], MA 73795!, MPU 007148 [photo!]*, S 08-16444 [photo!]) Notes:—Specimens have the indication of “cotypus” (= syntype) on the original label. Mateos & Valdés (2005) mention the BC specimen as original material, and Dobignard (2009) the MPU specimen as isotype. According to Mateos & Valdés (2005) the name should be attributed to Pau, since Font Quer would have merely transcribed a letter written by Pau. But Font Quer (1931c) explicitly indicates that he described this plant, therefore the authority should be “Pau ex Font Quer”, as stated by Paiva & Fernández Casas (2002) and Paiva (2015). The BM specimen was labelled as isotype and identified as P. rupestris subsp. fontqueri by Paiva in 1983.

1,2,3 Polygala sennenii Pau in Font Quer, Iter Marocc. 1929 nº 293 (1930a), in sched.

= Polygala baetica Willk.

Ind. loc.:—“Hab. in quercetis suberis, c. Asib de Ktama, ad 1600 m. alt.; 30 aprilis”

Lectotype (designated here):—In quercetis suberis, c. Asib de Ktama , 1600 m, 30 April 1929, Font Quer 293- 1929 (lectotype BC 826267 ! [https://www.ibb.csic.es/herbari/ JPEG / BC826267 .jpg]; isolectotypes BC 810596 !, BCN 72428 [photo!], GDA 35981 [photo!], MA 73951 !, MPU 006665 [photo!], RAB 061255 [photo!]*, S 08- 16001 [photo!])

Notes:—The name is cited as “ P. senneniana Pau ” in Font Quer (1930b). The RAB specimen was mentioned as type by El Oualidi et al. (2012). The GDA specimen was labelled as syntype by Vizoso in 2013. The MA specimen was identified as P. baetica by Paiva in 1999. The chosen lectotype is the BC specimen with more flowers.

1,2,3 Polygonum rhizoxylon Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 146 (1928), in sched.

≡ Polygonum balansae subsp. rhizoxylon (Pau & Font Quer) Greuter & Burdet

Ind. loc.:—“ Hab. in cacuminibus montis Iguermalez (Atlante rhiphaeo), solo schistoso, 1950 m. alt.; 26 junii” Lectotype (designated here):—In cacuminibus montis Iguermalez (Atlante rhiphaeo), 1950 m, 26 June 1927, Font Quer 146-1927 (lectotype BC 56280 ! [https://www.ibb.csic.es/herbari/ JPEG / BC56280 .jpg]; isolectotypes BCN 17959 [photo!], G 00437549 [photo!], GDA 36063 [photo!], MA 28176 !, MPU 006349 [photo!], S 07- 12657 [photo!])

Note:—The MA and GDA specimens were labelled as syntypes by Carrasco in 1974 and Vizoso in 2013, respectively, and the MPU as isotype by Sennedot and Scouppe in 2006.

1,2,3 Potentilla hirta var. afra Pau & Font Quer in Font Quer, Iter Marocc. 1929 nº 223 (1930a), in sched.

= Potentilla recta L.

Ind. loc.:—“Hab. in pratis montis Djebel Daddoh (Ktama), ad 1850 m. alt.”

Lectotype (designated here):—In pratis montis Djebel Daddoh (Ktama), 1850 m, 28 June 1929, Font Quer 223- 1929 (lectotype BC 98061 ! [https://www.ibb.csic.es/herbari/ JPEG / BC98061 .jpg]; isolectotypes BC 810651 !, BCN 72413 [photo!], GDA 38170 [photo!], MA 55456 !, MPU 006678 [photo!])

Notes:—This taxon was recorded as P. recta var. afra Pau & Font Quer by Carrasco (1975). The GDA specimen was labelled as syntype by Vizoso in 2013. The selected lectotype is the BC specimen that bears the most complete plant. The BC specimens were identified as P. recta by Romo in 1995.