Mannerheimia zanettii Shavrin, 2021

Shavrin, Alexey V., 2021, On the genus Mannerheimia Mäklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini), with taxonomic and faunistic notes on some species, Zootaxa 5040 (3), pp. 301-333 : 329-330

Mannerheimia zanettii Shavrin , sp.n.

( Figs. 9 , 40 , 70–71 )

Type material examined: Holotype ♂: ‘ Pakistan sett. | Hazara-Kaghan | Valley-Malkandi’ <printed>, ‘m 1350 | 20. 4. 1984 | leg. Zoia’ <printed>, ‘ HOLOTYPE | Mannerheimia | zanettii sp.n. | Shavrin A.V. des. 2019’ ( MCSNV).

Paratypes: 1 ♂ [dissected], 2 ♀♀ [one specimen without right antennomeres 8–11]: same data as the holotype , one specimens with label: ‘pr. [handwritten] | Mannerheimia [handwritten] | det. Zanetti 2009’ <printed>. All para- types with red printed label: ‘ PARATYPE | Mannerheimia | zanettii sp.n. | Shavrin A.V. des. 2019’ (1 ♀: cSh 1 ♂, 1 ♀: cZan).

Description. Measurements (n=4): HW: 0.54–0.67; HL: 0.37–0.42; OL: 0.18–0.20; AL (holotype): 1.07; PL: 0.49–0.52; PW: 0.91–0.95; EL: 1.22–1.30; EW: 1.20–1.41; AW: 1.26–1.39; MTbL (holotype): 0.65; MTrL (holotype): 0.43 (MTrL 1–4: 0.28; MTrL 5: 0.15); AedL: 0.50; TL: 2.60–2.90 (holotype).

Body moderately small and wide ( Fig. 9 ). Head and antennomeres 6–11 brown; pronotum, elytra and abdomen yellow to yellow-brown; mouthparts, antennomeres 1–5, legs and paratergites yellowish. Forebody shiny, without microsculpture; abdomen with fine transverse microreticulation. Head with sparse, moderately large and deep punctation, denser in middle; neck with large and deep punctures in middle; punctation of pronotum distinctly denser, larger and deeper than that on head, finer in middle and sometimes with impunctate portion in mediobasal third; scutellum with several moderately large and deep punctures; punctation of elytra about as that on pronotum, finer and coarser in prescutellar portion and finer in middle along suture, each elytron forming four to five longitudinal rows of punctures; abdominal tergites with fine and dense punctation.

Head 1.4–1.5 times as broad as long, somewhat flattened in middle, with moderately deep anteocellar foveae, convergent latero-anteriad toward level of middle length of eyes. Ocelli moderately large, situated about level of posterior margins of eyes; distance between ocelli slightly longer than distance between ocellus and posterior margin of eyes. Apical segment of maxillary palpi more than twice as long as preceding segment. Antennomere 3 distinctly narrower and slightly shorter than preceding antennomere, 4 1.5 times as long as broad, about as broad as 3, 5 slightly broader than 4, 6 broader and shorter than 5, 7–9 broader than 6, 10 about as long as broad, apical antennomere about 1.6 times as long as preapical antennomere.

Pronotum 1.8 times as broad as long, 1.4–1.6 times as broad as head, widest in middle, more narrowed posteriad than anteriad; apical margin rounded, about as long as rounded posterior margin; anterior angles widely rounded, not protruded anteriad; posterior angles rounded, with slightly sinuate laterobasal margins in front of it; laterobasal portions distinctly depressed.

Elytra about as long as broad or slightly broader than long, more than twice as long as pronotum, gradually widened apicad.

Metatarsus 1.5 times as long as metatibia.

Male. Apical margins of abdominal tergite VIII deeply and abdominal sternite VIII slightly concave. Aedeagus with wide basal portion, significantly narrowed toward subacute apex; parameres moderately wide, slightly exceeding apex of median lobe; internal sac with two moderately wide and long fields of elongate thorns ( Fig. 70 ). Lateral aspect of aedeagus as in Fig. 71 .

Female. Apical margin of abdominal tergite VIII straight or rounded. Apical margin of abdominal sternite VIII widely rounded.

Comparative notes. Regarding the general shape of the aedeagus, M. zanetti is similar to M. cachemirica and M. vicina from Kashmir, but distinguished by paler coloration, a more transverse pronotum with the anterior angles not protruding anteriad, and a broader median lobe and broader apical portions of the parameres.

Distribution. The new species is at present known only from the type locality ( Fig. 40 ) in Hazara Kaghan Valley, North Pakistan.

Bionomics. The specimens were collected at an elevation of 1350 m a.s.l. Detailed ecological data are unknown.

Etymology. Patronymic, the species is named to honour my colleague Adriano Zanetti (Verona).

Figures

FIGURES 1–9. Habitus of Mannerheimia: 1—M. afghanica (paratype), 2—M. arctica, 3—M. asiatica, 4—M. brevipennis, 5—M. japonica, 6—M. obscura, 7—M. pleshanovi (paratype), 8—M. yasudai, 9—M. zanettii (paratype). Scale bar: 1.0 mm.

FIGURES 39–40. Distribution of Mannerheimia arctica (Fig. 39) and M. brevipennis (Fig. 40). Black circles: material studied by authors, white circles: previously published material not seen by author.

FIGURES 70–71. Aedeagus of Mannerheimia zanettii (paratype): 70—parameral view, 71—lateral view. Scale bar: 0.1 mm.