Isopedhispa costata, Borowiec & Świętojańska & Sekerka, 2019

Borowiec, Lech, Świętojańska, Jolanta & Sekerka, Lukáš, 2019, Revision of the tribe Cryptonychini (Coleoptera: Chrysomelidae: Cassidinae) of New Caledonia, Zootaxa 4690 (1), pp. 1-71 : 31-33

Isopedhispa costata sp. nov.

( Figs 101–107 , 226)

Etymology. Named after costate elytra.

Type locality. New Caledonia, Mt. Ignambi , 900 m a.s.l.

Diagnosis. Isopedhispa costata sp. nov. with I. latemarginata sp. nov. are the only species with body mostly black and pronotum coarsely and densely punctate on almost entire dorsal surface and sides of pronotum not or very shallowly concave behind anterolateral projections while Isopedhispa cocotis ( Maulik, 1933) and Isopedhispa ferruginea Spaeth, 1936 have dorsum from ferrugineous to brown, pronotum with large impunctate areas and sides of pronotum behind anterolateral projections distinctly concave. Isopedhispa costata and I. latemarginata at first glance appear similar but I. costata differs in larger size of body with length 11.45–12.95 mm (vs. 6.60–9.50 mm in I. latemarginata ), elytra uniformly black including explanate margin ( I. latemarginata has explanate margin from ferrugineous to ferrugineous-brown), and elytra more costate with elytral interval 5 costate on whole length, interval 7 costate behind the middle, and interval 9 costate in anterior half (in I. latemarginata only interval 7 is elevated behind middle and interval 9 more or less elevated on anterior half).

Description. Length 11.45–12.95 mm, width 4.10–4.90 mm.

Entire dorsum, antennae and legs black ( Fig. 101 ). Thorax black or with central part of expanded apex of prosternal process, mesoventrite and intercoxal process of metaventrite ferrugineous-brown to brown. Abdomen black, ventrites laterally with small ferrugineous-brown patches of diffused borders. Body glabrous except for yellowish hairs on frontoclypeus, short golden brown pubescence on distal antennomeres and golden orange pubescence on tarsal pads and apices of tibiae. Head transverse 2.1–2.2 × as wide as long, interocular plate rectangular, convex but dorsal surface almost flat, separated from vertex by deep sulcus, anterior corners of interocular plate forming sharp denticles, along anterior and lateral margins run distinct sulci. Surface of interocular plate coarsely and densely punctate, appears rugose, with very deep median sulcus along whole length; interantennal process short, approximately as long as ¼ length of antennomere I, not sexually dimorphic, flattened laterally, keel-shaped, acute apically, without sulcus dorsally ( Fig. 104 ). Frontoclypeus moderately elongate, 1.5 × as long as broad (including interantennal process), acute apically, anterior corners forming straight angles, surface on basal half with moderately coarse setose punctures, in area close to anterior angles flat, impunctate or with 1–2 moderately coarse punctures, along middle runs narrow median keel ( Fig. 103 ). Antennae 0.33 × as long as body, slightly compressed apically; antennomere I large, approximately 1.6 × as long as broad; antennomere II short, 1.2–1.3 × as long as broad, approximately 0.4 × as long as I; antennomeres III–VII elongate, 1.4–1.6 × as long as II; antennomeres VIII–X slightly longer than II; antennomere XI 2.1 × as long as X, subangulate apically ( Fig. 102 ). Pronotum approximately 1.1 × as broad as long, subpentagonal, in anterolateral parts with small projections, sides behind projections shallowly concave, diverging posteriorly, anterior margin strongly convex, basal margin bisinuate; sharp lateral margins forming narrow explanation along entire length, interrupted only by narrow emargination in front of basal tubercles, anterior angles obtuse with sharp anterior tubercle, basal angles bearing small, sharp tooth ( Fig. 105 ); disc on sides regularly convex, top flattened, without impressions or with shallow round impression in front of scutellum and two round to transverse ones, shallow impressions in middle of pronotum, without medial sulcus, whole surface with coarse and dense punctures, interspaces narrower than puncture diameter and surface appears partly irregular. Elytra twice as long as broad, subparallel-sided basally and distinctly broadened posteriorly, widest somewhat behind middle, and lateral sides with broadly explanate and impunctate margins. Apex of elytra regularly rounded, with obtuse sutural angle and broadly rounded lateral angle. Disc with 8–11 punctures in scutellar row; eight rows in posthumeral part, ten rows behind middle and ten rows apically but lateral interspaces apically with several additional irregular punctures thus regularity of rows partly disturbed; intervals 1–4 flat, as wide as or slightly wider than rows, interval 5 forming sharp costa on almost whole length, interval 7 elevated and partly costate apically and humeral interval costate in anterior half and obtusely elevated in posterior half, intervals 3, 5, 7, 9 more or less distinctly elevated on slope. Ventral surface mostly shiny, hypomera coarsely but moderately densely punctate; prosternum smooth and shiny, between coxae as wide as ⅓ width of coxa, expanded apex shallowly impressed centrally, anterior margin of prosternum elevated, separated from prosternal process by row of punctures, mesoventrite mostly smooth and shiny with indistinct oblique striation, metaventrite laterally coarsely and densely punctate, central part and anterior process mostly with fine longitudinal and oblique striation; abdomen mostly smooth and shiny with very fine pricks. Legs stout ( Fig. 106, 107 ), sexually dimorphic.

Sexual dimorphism distinct, in males ventral side of mid femora with angulation in middle, mid tibiae before apex curved and enlarged similarly as in I. latemarginata (Fig. 126), with distinct spine apically, last ventrite deeply emarginate; in females mid femora without angulation on ventral side, mid tibiae stout, straight, regularly broadened from base to ⅔ length, with ventral side regularly convex, apex with minute spine, last ventrite regularly shallowly emarginate apically.

Type material. Holotype: ♂, ‘ NEW CALEDONIA | Mt. Ignambi , 900 m | 4.II.1964 [w, p, cb] || swept [w, p, cb] || R. Straatman | Collector | BISHOP [w, p, cb]’ ( BPBM) . Paratypes: 1 ♀, ‘ NEW CALEDONIA (N) | -20.5702 / 164.7739 | Mt. Paniè E trail | 900-1300 m rainforest | 23.11.210 | leg. R. Ruta, M. Wanat [w, p, cb]’ ( WMNH) ; 1 ♀, ‘ NEW CALEDONIA | Mt. Panier , 500- | 1000m, 11.X.1967 | J. Sedlacek [w, p, cb] || J. & M. Sed- lacek | Collectors | BISHOP [w, p, cb] || M.T. 7. [w, hw, s]’ ( LS) .

Figures

FIGURES 101–107. Isopedhispa costata sp. nov. 101. Body dorsal, 102. Antenna, 103. Frontoclypeus and prosternum, 104. Head dorsal, 105. Pronotum, 106. Fore leg of female, 107. Mid leg of female.