Commatarcha Meyrick, 1935

Qian, Shuonan & Li, Houhun, 2018, Taxonomic review of the genus Commatarcha Meyrick (Lepidoptera: Carposinoidea: Carposinidae) from China, with descriptions of four new species, Zootaxa 4418 (5), pp. 432-448 : 432-433

Commatarcha Meyrick, 1935

Commatarcha Meyrick, 1935a: 594 . Type species: Commatarcha palaeosema Meyrick, 1935 .

Delarchis Meyrick, 1938: 15 . Type species: Delarchis citrogramma Meyrick, 1938 .

Generic characters. Adult ( Figs 2 ¯4). Head with appressed scales. Proboscis well-developed. Labial palpus upturned in male ( Fig. 2a ), only extended forward and longer in female ( Fig. 2b ); second segment usually longer than diameter of eye; third segment short, greyish, white at apex. Antenna filiform, cilia in male usually same length as diameter of flagellum. Forewing narrowly elongate, with six or more scale tufts roughly scattered in discal cell, in some species often with a triangular or sub-triangular black basal patch ( Fig. 3 ); hindwing trapezoid. Wing venation ( Fig. 4 ): forewing with R veins separate, M1 closer to R5 than to M2, M3 and CuA1 close at base and from lower angle of cell, CuA2 from lower margin before lower angle of cell, 1A+2A furcate basally; hindwing with Rs reaching apex, M1 developed or fully vestigial, M2 absent, M3 and CuA1 from lower angle of cell, stalked at base, CuA2 from about distal 1/4 of lower margin of cell.

Male genitalia ( Fig. 5 ). Uncus reduced to a small conical lobe, fused to tegumen. Gnathos reduced. Tegumen short, narrowly banded anterolaterally. Valva varied in shape, with a rod-shaped, weakly sclerotized dorsobasal process and a wedge-shaped, heavily sclerotized ventrobasal process, both processes usually extending outward parallelly. Transtilla usually banded. Vinculum elongate triangular, narrowed anteriorly; saccus slender, narrowly rounded or pointed apically. Juxta often short. Ectophallus (specialized anellus) often developed, triangular. Aedeagus elongate, slender; cornutus being a single spine, or double spines, or a cluster of fine spines.

Female genitalia ( Fig. 6 ). Eighth sternum weakly sclerotized. Ostium bursae large and elongate. Antrum (= colliculum in Diakonoff 1989) usually well-developed, heavily sclerotized. Ductus bursae slender. Corpus bursae elongate ovate; signum absent.

Diagnosis. Commatarcha is similar to Bondia Newman, 1856 by the forewing having several scale tufts. It can be distinguished from Bondia by male genitalia: the valva has two dorsobasal processes in Commatarcha ; Bondia has a valva with distal half often divided into two to four lobes, but lacking the dorsobasal processes ( Davis 1969).

Distribution. China, Japan, Kashmir, Korea, Pakistan.

Figures

FIGURES2‒6. Genericcharacters of Commatarchaspp.2, Head of C. angustiptera Li showinglabial palpus:2a, ♂; 2b, ♀; 3, Wing venation of C. palaeosema Meyrick, ♂, slide No. QSN17502; 4, Wing venation of C. convoluta sp. nov., ♀, slide No. QSN17477W; 5, Male genitalia of C. setiferaedeaga sp. nov., paratype, slide No. QSN17487; 6, Female genitalia of C. convoluta sp. nov., paratype, slide No. QSN17477 (Scale bars = 0.5 mm).