Integrivalvia, Volynkin & Huang, 2019

Volynkin, Anton V., Huang, Si-Yao & Ivanova, Maria S., 2019, An overview of genera and subgenera of the Asura / Miltochrista generic complex (Lepidoptera, Erebidae, Arctiinae). Part 1. Barsine Walker, 1854 sensu lato, Asura Walker, 1854 and related genera, with descriptions of twenty new genera, ten new subgenera and a check list of taxa of the Asura / Miltochrista generic complex, Ecologica Montenegrina 26, pp. 14-92 : 66

Genus Integrivalvia Volynkin & Huang, gen. nov.

https://zoobank.org/ urn:lsid:zoobank.org:act:AB544F1E-96AD-42FC-8955-63BCAF1C3DD4 ( Figs 98, 99 , 156 , 208 )

Type species: Barsine exclusa Butler, 1877 .

Etymology. The generic name refers to its entire valvae without any costal and saccular processes. Gender feminine.

Diagnosis. Small moths with reddish or orange coloration and blackish or brown pattern. The male genital capsule is characterised by the entire, narrow, lobe-like valva without any processes (similar to that of the African genus Tumicla and the Indian Parvuspina only). The autapomorphic feature of the new genus is the presence of a very long and narrow granulated distal diverticulum of aedeagus vesica having a hook-like curved robust plate at base (unlike Hampsonascia , which has long and narrow diverticula also, but without plates at base). In addition, the aedeagus vesica has several short diverticula bearing fields of granulation and patches of spinules. The basal plate of vesica ejaculatorius is absent. The autapomorphic feature in the female genitalia is the presence of a membranous lateral semiglobular protrusion on the connection of the ductus bursae with the corpus bursae. In addition, the female genitalia of Integrivalvia are characterised by the combination of the following features: (1) the sclerotised, dorso-ventrally flattened ductus bursae without antrum; (2) the pear-shaped corpus bursae densely covered with small spinulose scobination and having a band-like cluster of large spinules medially and a broad field of large spinules laterally and at the base of appendix bursae; (3) the appendix bursae is directed posteriorly, elongated, conical, curved, densely covered with robust spinules.

Distribution. The genus is widespread from southern China (Hainan) through eastern Indochina and Malay Peninsula to Greater and Lesser Sunda Islands including the Philippine Palawan Island.

Number of species. The genus comprises its type species with one subspecies. In addition, herein we provisionally include to the genus seven New Guinean species and one subspecies treated by de Vos (2019) to be members of the genus Asuridia and, in addition, one New Guinean species previously included to the genus Melanaema . However, all those New Guinean taxa are different in their genitalia structure from the type species (e.g., the principally different vesica structure having no characteristic elongated diverticulum with a sclerotised plate at base) and their generic placement needs further clarification.

Figures

Figures 82–103. Type species of genera and subgenera of the Asura / Miltochrista generic complex: adults. Depositories of specimens: 82, 84, 85, 90–93, 98, 99 – NHMUK; 83, 88, 89, 97, 100, 101 – MWM/ZSM; 86, 87, 94, 95 – CKC; 96, 102, 103 – ZFMK.

Figures 156–159. Members of the Asura / Miltochrista generic complex: male genitalia. 156–158 – type species of genera.

Figures 205–211. Type species of genera of the Asura / Miltochrista generic complex: female genitalia.