Pinobius forcipifer, Assing, 2018

Assing, Volker, 2018, A revision of Pinobius. II. Three new species from Cambodia and Indonesia, and additional records (Coleoptera: Staphylinidae: Paederinae), Linzer biologische Beiträge 50 (2), pp. 997-1004 : 1003-1004

Pinobius forcipifer nov.sp. ( Figs 9-15 )

T y p e m a t e r i a l: Holotype ♂: " Indonesia, E Kalimantan, ca. 55 km W of Balikpapan, PT Fajar Surya Swadaya (area), 01°18.3'S, 116°21.0'E, 100 m, J. Hájek, J. Schneider & P. Votruba leg. / border of Acacia mangium plantation and primary rainforest, stream and waterfal [sic], puddles, individually collecting, + light trap / Holotypus ♂ Pinobius forcipifer sp. n., det. V. Assing 2015" ( NMP). Paratypes: 2♂♂, 2♀♀, same data as holotype ( NMP, cAss).

E t y m o l o g y: The specific epithet (from forceps: pliers, forceps) is an adjective alluding to the shape of the apex of the dorsal plate.

D e s c r i p t i o n: Body length 9.5-10.0 mm; length of forebody 4.9-5.4 mm. Habitus as in Fig. 9 . Coloration: head and pronotum dark-brown; elytra reddish to dark-reddish, with the anterior and lateral portions darker; abdomen dark-brown with reddish apex (segments VIII-X); legs pale-brown; antennae dark-reddish.

Head ( Fig. 10 ) moderately transverse, approximately 1.15 times as broad as long; lateral margins behind eyes subparallel in dorsal view; punctures of variable sizes and moderately dense, somewhat sparser in median dorsal portion; interstices without microsculpture, except for indistinct micropunctures. Eyes weakly convex and approximately as long as postocular region in dorsal view. Antenna ( Fig. 11 ) 2.6-2.8 mm long; antennomeres IV-VI weakly oblong; VII approximately as long as broad; VIII-X weakly transverse.

Pronotum ( Fig. 10 ) weakly oblong, approximately 1.05 times as long as broad and as broad as head; punctation similar to that of head; interstices without microsculpture; midline moderately broadly impunctate.

Elytra ( Fig. 10 ) nearly as long as pronotum; punctation defined, denser than that of pronotum. Hind wings fully developed. Metatarsomere I slightly longer than the combined length of metatarsomeres II and III.

Abdomen narrower than elytra; punctation fine and rather dense; interstices with shallow microreticulation; posterior margin of tergite VII with palisade fringe; posterior margin of tergite VIII strongly convex posteriorly.

♂: protarsomeres I-IV strongly dilated; sternite VII not distinctly modified; sternite VIII ( Fig. 12 ) approximately as broad as long, posterior incision narrow and deep, its depth approximately 0.6 times the length of sternite; aedeagus ( Figs 13-15 ) 1.6-1.7 mm long (measured from apex of dorsal plate), dorso-ventrally flattened, and asymmetric; ventral process apically obliquely bisinuate in ventral view; dorsal plate apically with a conspicuous and asymmetric bifid process; parameres slender and of subequal length, apically extending approximately to apex of dorsal plate, left paramere (ventral view) somewhat dilated apically; internal sac with a long, rather stout, strongly sclerotized, and somewhat S-shaped apical structure on the left (ventral view) and a slightly shorter, more slender, weakly sclerotized, and simply curved structure in more basal position on the right.

♀: protarsomeres I-IV strongly dilated, only slightly less so than in male.

C o m p a r a t i v e n o t e s: Pinobius forcipifer belongs to the P. major group and is distinguished from all its congeners particularly by the conspicuous shape of the dorsal plate of the aedeagus. Regarding its male sexual characters, it is most similar to P. firmilobatus ASSING, 2014 (Southeast Asia from Thailand to Borneo) and P. baculatus ASSING, 2014 (Java, Sumba), from which it additionally differs by a slightly less robust habitus with a less transverse head and a more distinctly oblong pronotum, by relatively shorter, more slender, less strongly sclerotized, and differently shaped parameres (P. firmilobatus), and by a deeper posterior incision of the male sternite VIII (P. baculatus).

D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: Thetypelocalityissituated in Kalimantan Timur, southeastern Borneo. The specimens were collected at the border between an Acacia mangium plantation and a primary rainforest at an altitude of 100 m.

Figures

Figs 9-15: Pinobius forcipifer nov.sp.: (9) habitus; (10) forebody; (11) antenna; (12) male sternite VIII; (13-14) aedeagus in lateral and in ventral view; (15) aedeagus in transparent light in ventral view. Scale bars: 9: 2.0 mm; 10-11: 1.0 mm; 12-15: 0.5 mm.